[HOME] [INDEXES] < Life < Insecta < Lepidoptera < Ditrysia < Papilionoidea < Pieridae < Pierinae
Belenois Hübner, [1819]
Caper Whites
Belenois Hübner, [1819]; Verz. bek. Schmett. (6): 92, TS: Papilio calypso Drury
Anaphaeis Hübner, [1819]; Verz. bek. Schmett. (6): 93, TS: Papilio creona Cramer
Glycestha Billberg, 1820; Enum. Ins. Mus. Billb.: 76, TS: Papilio coronea Cramer
Pseudohuphina Stoneham, 1940; Bull. Stoneham Mus. (40): 4, TS: Pieris raffrayi Oberthür
Pseudanaphaeis Bernardi, 1953; Rev. franc. Ent. 20 (1): 50, TS: Pieris gidica Godart
Belenois, NSG Voucher Specimen [Wahlberg]
Belenois, Pieridae Holarcticae [Heiner Ziegler]
Annaphaeis ; Butler, 1870, (missp.)
Anapheis[sic] ; [BAUR, 153 (missp)]
Belenois ; Winhard, 2000, Butterflies of the world, 10: 27; [AfrL]

The "pseudohuphina" species-group

Belenois raffrayi (Oberthür, 1878)
Ethiopia, Sudan, W.Kenya, Zaire, Rwanda, W.Tanzania. See [About maps] Raffray's White
Pieris raffrayi Oberthür, 1878; Etud. Ent. 3: 17, pl. 1, f. 3
Pseudohuphina raffrayi ;
Belenois raffrayi ; [EBW]; [AfrL]

Larva on Capparis, Rhus [BK]

S.Ethiopia, S.Sudan Belenois raffrayi raffrayi ; [BAFR, 76 (text)]; [AfrL]
B. r. extendens (Joicey & Talbot, 1927)
W.Kenya, Uganda, Rwanda, Burundi, E.Zaire, W.Tanzania? Belenois raffrayi extendens ; [BAFR, 76]; [BK, 138]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 44, f. 12; [AfrL]
B. r. similis Kielland, 1978
W.Tanzania Belenois raffrayi similis ; [AfrL]

Belenois margaritacea Sharpe, 1891
Kenya, Tanzania. See [About maps] Margarita's Caper White
Pseudohuphina margaritacea ;
Belenois margaritacea ; [BK, 138]; [AfrL]

Larva on Capparis, Maerua, Ritchiea [BK]

SW.Kenya (Sotik) Belenois margaritacea margaritacea ; [BAFR, 76]; [BK, 138 (note)]; [AfrL]
B. m. kenyensis (Joicey & Talbot, 1927)
SE.Kenya (Teita Hills, Chyulu Hills) Belenois margaritacea kenyensis ; [BAFR, 76]; [BK, 138 (note)]; [AfrL]
B. m. plutonica (Joicey & Talbot, 1927)
N.Tanzania (Mt. Kilimanjaro, Usambara Mts., Ngorongoro Crater) Belenois margaritacea plutonica ; [BAFR, 76]; [AfrL]
B. m. somereni (Talbot, 1928)
Kenya (Meru, Matthew's Range) Belenois margaritacea somereni ; [BK, 138 (note)]; [AfrL]
B. m. intermedia Kielland, 1982
Belenois margaritacea intermedia ; [AfrL]

The ? species-group

Belenois albomaculatus (Goeze, 1779)
Belenois albomaculatus ; [AfrL]

Belenois grandidieri (Mabille, 1878)
Madagascar, Aldabra Is.. See [About maps] Pieris grandidieri Mabille, 1878; , TL: Madagascar
Belenois grandidieri ; [BAFR, 76]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 6; [AfrL]
Belenois (Belenois) grandidieri ; [MadL]

Belenois larima (Boisduval, 1836)
Belenois larima ; [AfrL]

Belenois zochalia (Boisduval, 1836)
Cape - Moçambique, Rhodesia - Ethiopia, Uganda. See [About maps] Forest White Forest Caper White
Pieris zochalia Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1: 506
Belenois zochalia ; [AfrL]

Larva on Niebuhria pedunculosa, Maerua triphylla [PBSA]

South Africa, Rhodesia Belenois zochalia zochalia ; [BAFR, 76]; [AfrL]
B. z. agrippinides (Holland, 1896)
Malawi, Tanzania, Kenya, Uganda Belenois zochalia agrippinides ; [BAFR, 76]; [BK, 139]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 12; [AfrL]

Larva on Capparidaceae, Brassica, Salvadora [BK]

B. z. connexiva (Joicey & Talbot, 1927)
Cameroun (highlands) Belenois zochalia connexiva ; [BAFR, 76 (text)]; [AfrL]
B. z. galla (Ungemach, 1932)
Ethiopia Belenois zochalia galla ; [BAFR, 76 (text)]; [AfrL]
?B. z. camerounica Bernardi, 1966²
Nigeria - Cameroun (hihlands) ?Belenois zochalia camerounica ; [BK, 139 (note)]

Belenois ?sp.
Belenois ?sp. ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 13 (Mpanda, Ntakatta, Kigoma, Tanzania)

Belenois crawshayi Butler, 1894
Angola, Zambia - E.Zaire, Uganda, W.Kenya, Tanzania. See [About maps] Crawshay's Caper White
Belenois crawshayi ; [BAFR, 78]; [BK, 139]; [AfrL]

Larva on Capparis [BK]


Belenois diminuta Butler, 1894
Tanzania. See [About maps] Belenois diminuta Butler, 1894; Proc. zool. Soc. Lond.: 666, pl. 60, f. 7
Belenois diminuta ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 14; [AfrL]

Belenois victoria Dixey, 1915
Uganda, Burundi, Rwanda, Tanzania, E.Zaire, W.Kenya. See [About maps] Victoria White
Belenois victoria Dixey, 1915; Trans. Ent. Soc.: 4, pl. 1, f. 5-6
Belenois victoria ; [AfrL]

Larva on Capparis, Maerua, Ritchiea [BK]

W.Kenya (highlands, W of Rift Valley), Uganda Belenois victoria victoria ; [BK, 139]; [BAFR, 78]; [AfrL]
B. v. hecqi Berger, 1953
NE.Zaire Belenois victoria hecqi ; [BAFR, 78 (text)]; [AfrL]
B. v. schoutedeni Berger, 1953
W.Uganda, E.Zaire (Kivu) Belenois victoria schoutedeni ; [BAFR, 78]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 15, 18; [AfrL]

The ? species-group

Belenois subeida (C. & R. Felder, 1865)
Guinea - Ethiopia, W.Kenya, Tanzania. See [About maps] Belenois subeida ; [AfrL]
B. s. frobeniusi (Strand, 1909)
W.Africa Belenois subeida frobeniusi ; [BAFR, 80 (text)]; [AfrL]
Ethiopia Belenois subeida hiemalis ; [BAFR, 80 (text)]
B. s. sylvander Grose-Smith, 1890
W.Kenya, Uganda, Tanzania, N.Zaire, C.A.R. Belenois subeida sylvander ; [BAFR, 80]; [BK, 140]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 9; [AfrL]

Larva on Capparis, Maerua, Ritchiea [BK]

B. s. hailo (Ungemach, 1932)
S.Ethiopia, SE.Sudan Belenois calypso hailo ; [BAFR, 82 (text)]
Belenois subeida hailo ; [AfrL]

Belenois calypso (Drury, [1773])
Gambia - Angola, W.Kenya, Tanzania. See [About maps] Calypso Caper White
Papilio calypso Drury, [1773]; Illust. Nat. Hist. Exot. Insects 2: index, 30, pl. 17, f. 3-4
Belenois calypso ; [EBW]; [AfrL]
Nigeria - Ghana, Ivory Coast, Liberia, Sierra Leone Belenois calypso calypso ; [BAFR, 82]; [AfrL]
B. c. dentigera Butler, 1888
Cameroun - Zaire, NW.Angola Belenois calypso dentigera ; [BAFR, 82 (text)]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 5-6; [AfrL]
B. c. minor Talbot, 1943
Uganda, W.Kenya (W of Rift Valley) Belenois calypso minor ; [BAFR, 82 (text)]; [BK, 140]; [AfrL]

Larva on Capparis, Cadaba, Maerua, Ritchiea [BK]

B. c. marlieri Berger, 1981
Belenois calypso marlieri ; [AfrL]

Belenois welwitschii Rogenhofer, 1890
-. See [About maps] Belenois welwitschii ; [AfrL]
NE.Angola, S.Zaire (Shaba), Zambia, Malawi, S.Tanzania, W.Tanzania Belenois calypso welwitschii ; [BAFR, 82]
Belenois welwitschii welwitschii ; [AfrL]
B. w. shaba Berger, 1918
Belenois calypso shaba Berger, 1981
Belenois calypso shaba ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 7-10
Belenois welwitschii shaba ; [AfrL]

Belenois sudanensis (Talbot, 1929)
Cameroun - W.Kenya, Tanzania. See [About maps] Sudan Caper White
Belenois sudanensis ; [BK, 140]; [AfrL]

Larva on Clerodendrum [BK]

B. s. sudanensis (Talbot, 1929)
Belenois sudanensis sudanensis ; [AfrL]
Belenois sudanensis pseudodentigera ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 12; [AfrL]
B. s. katalensis Berger, 1981
Belenois sudanensis katalensis ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 15; [AfrL]
B. s. mayumbana Berger, 1981
Belenois sudanensis mayumbana ; [AfrL]

Belenois rubrosignata (Weymer, 1901)
Cameroun - S.Sudan, Uganda, W.Kenya, Tanzania. See [About maps] Red-Edged White
Belenois rubrosignata ; [EBW]; [AfrL]
Angola, S.Zaire (Shaba), Zambia Belenois rubrosignata rubrosignata ; [BAFR, 80]; [BK, 141]; [AfrL]
B. r. kongwana Talbot, 1943
Malawi, Tanzania, S.Uganda Belenois rubrosignata kongwana ; [BAFR, 80 (text)]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 44, f. 2-3; [AfrL]
B. r. peeli Dixey, 1900
Kenya, Somalia Belenois rubrosignata peeli ; [AfrL]; [BAFR, 80 (text)]

Belenois thysa (Hopffer, 1855)
Natal, Moçambique, Rhodesia - Kenya, Ethiopia, Somalia. See [About maps] False Dotted Border
Pieris thysa Hopffer, 1855; Mber. dt. Akad. Wiss. Berl. 1855: 639
Pieris thysa malaria Suffert
thysa f. malaria ; [wet season f.]
Belenois thysa ; [EBW]; [BK, 141]; [AfrL]

Larva on Capparis sp. [PBSA], Maerua, Ritchiea [BK]

E.Cape - Natal, Mozambique, Rhodesia, Zambia, Malawi, Tanzania, Kenya (coast) Belenois thysa thysa ; [BAFR, 82]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 16, pl. 44, f. 1, back f. 5; [AfrL]
B. t. tricolor Talbot, 1943
Ethiopia Belenois thysa tricolor ; [BAFR, 82]; [AfrL]
B. t. meldolae Butler, 1872
Angola, S.Zaire - Uganda, W.Kenya, S.Sudan Belenois thysa meldolae ; [BAFR, 82 (text)]; [AfrL]

Belenois solilucis Butler, 1874
S.Ethiopia, S.Sudan - Uganda, NW.Tanzania, W.Kenya, Cameroun, Angola. See [About maps] Belenois solilucis ; [EBW]; [BK, 141]; [AfrL]
N.Angola, Zaire - W.Tanzania, Cameroun Belenois solilucis solilucis ; [BAFR, 85 (text)]; [AfrL]
B. s. loveni (Aurivillius, 1921)
Uganda, NE.Zaire, S.Sudan, SW.Ethiopia Belenois solilucis loveni ; [BAFR, 85]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 44, f. 10-11; [AfrL]

Belenois aldabrensis (Holland, 1896)
Aldabra, Astove, Assumption Is. Belenois aldabrensis ; [AfrL]; [BAFR, 85]

Belenois anomala (Butler, 1881)
Socotra I. Belenois anomalis ; [BAFR, 86]
Belenois anomala ; [AfrL]

?Belenois ogygia (Trimen, 1883)
Natal. See [About maps] Pieris ogygia Trimen, 1883; Trans. ent. Soc. Lond. 1883: 356
Belenois ogygia ; [BAFR, 79]

The ? species-group

Belenois antsianaka (Ward, 1870)
Madagascar. See [About maps] Pieris antsianaka Ward, 1870; Ent. mon. Mag. 7: 30, TL: Madagascar
ramona Grose-Smith²
Belenois antsiaka[sic?] ; [EBW]; [BAFR, 86]
Belenois antsianaka ; Winhard, 2000, Butterflies of the world, 10: 28, pl. 44, f. 4; [AfrL]
Belenois (Belenois) antsianaka ; [MadL]

Belenois mabella Grose-Smith, 1891
Madagascar. See [About maps] Belenois mabella ; [BAFR, 86]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 44, f. 9; [AfrL]
Belenois (Pseudanaphaeis) mabella ; [MadL]

Belenois java (Linnaeus, 1768)
Indonesia - Samoa, Australia. See [About maps] Caper White
Papilio java Linnaeus, 1768; in Sparrman, Amoenit. acad., 7 (150): 504, nota
Papilio coronea Cramer, [1775]; Uitl. Kapellen 1 (1-7): 106, pl. 68, f. B, C
Papilio clytie Donovan, 1805
Papilio deiopea Donovan, 1805
Pieris niseia Macleay, 1826
Anaphaeis nigrita Joicey & Talbot, 1928
Belenois java, NSG Voucher Specimen [Wahlberg]
Belenois java, Lepidoptera Larvae of Australia [Don Herbison-Evans]
Anaphaeis java ; [BOW: pl. 156, f. 11]
Anapheis java ; [EBW]; [BOR, 144]

Larva on Capparis mitchelli, C. nummularia, Apophyllum anomalum [Don Herbison-Evans], Capparis nobilis, C. lasiantha [BAUR], Apophyllum anomalum, Capparis arborea, C. lasiantha, C. mitchellii, C. nobilis, C. spinosa var. nummularia, C. canescens?, C. spp. K. L. & L. E. Dunn, 1991, Review Austr. Butts. (1-4): (120-140)

Java, Kangean - Sumbawa, Sumba, Sulawesi Anapheis java java
Anapheis java java ; [BOR, 144]
Belenois java java ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 10
Belenois java ?ssp. ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 11 (Gaibobo I., Papua New Guinea)
B. j. savuana Fruhstorfer²
Sawu I. (Savu) Anapheis java savuana ; [BOR, 144]
B. j. teutonia (Fabricius, 1775)
Australia Papilio teutonia Fabricius, 1775
Anapheis java teutonia ; [BAUR, 153]
B. j. ina Fruhstorfer²
Timor?, Wetar, Batar, Kissar ?anita Fruhstorfer
Anapheis java ina ; [BAUR, 153 (text)]
B. j. peristhene (Boisduval, 1859)
New Hebrides, New Caledonia, Loyalty Is., Norfolk I., Woodlark I. Pieris peristhene Boisduval, 1859
Anapheis java peristhene ; [BAUR, 153 (text)]
New Britain, New Ireland, Duke of York I. Anapheis java picata ; [BAUR, 153 (text)]
B. j. micronesia Fruhstorfer²
Fiji ?vitiensis Fruhstorfer
Anapheis java micronesia ; [BAUR, 153 (text)]

Belenois helcida (Boisduval, 1833)
Madagascar. See [About maps] Pieris helcida Boisduval, 1833; Faun. ent. Madagascar, Lépid.: 17, pl. 2, f. 1-2, TL: Madagascar
Belenois helcida ; [BAFR, 84]; [AfrL]
Belenois (Belenois) helcida ; [MadL]

Belenois hedyle (Cramer, 1777)
-. See [About maps] Papilio hedyle Cramer, 1777
Belenois hedyle ; [AfrL]
Sierra Leone, Liberia, Ivory Coast, Ghana, Benin Belenois hedyle hedyle ; [BAFR, 84 (text)]; [AfrL]
B. h. rhena (Doubleday, 1846)
Ghana - W.Nigeria Belenois hedyle rhena ; [BAFR, 84]; [AfrL]
B. h. ianthe (Doubleday, 1842)
Sierra Leone, Togo, Guinea, Guinea-Bissau Belenois ianthe ; [BAFR, 84]
Belenois hedyle ianthe ; [AfrL]

Belenois theora (Doubleday, 1846)
-. See [About maps] Belenois theora ; [EBW]; [AfrL]
Sierra Leone - Nigeria Belenois theora theora ; [BAFR, 81]; [AfrL]
?B. t. concolor Aurivillius²
Cameroun - C.A.R., C.Zaire, S.Zaire ?Belenois theora concolor ; [BAFR, 81 (text)]
B. t. ratheo (Suffert, 1904)
C.A.R Pieris theora ratheo Suffert, 1904; Dt. Ent. Z. Iris 17: 81
Belenois theora ratheo ; Winhard, 2000, Butterflies of the world, 10: 28, pl. 44, f. 7-8; [AfrL]
B. t. laeta (Weymer, 1903)
W.Uganda, E.Zaire, S.Sudan Belenois theora laeta ; [BAFR, 81 (text)]; [AfrL]

Belenois theuszi (Dewitz, 1889)
S.Angola, NW.Zambia, S.Zaire - Gabon, Cameroun, W.Nigeria. See [About maps] Belenois theuszi ; [BAFR, 82]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 43, f. 17; [AfrL]

Subgenus Anaphaeis Hübner, [1819]
Anaphaeis Hübner, [1819]; Verz. bek. Schmett. (6): 93

Belenois aurota (Fabricius, 1793)
Ceylon, India, NW.India, Pakistan, Afghanistan, Baluchistan, Arabia, SAF, Tropical Africa (dry zones). See [About maps] Brown-veined White Caper White Pioneer
Papilio aurota Fabricius, 1793; Ent. Syst. 3 (1): 197, TL: Tranquebar, S.India
Papilio mesentina Cramer, [1780]; Uitl. Kapellen 3 (23-24): 140, pl. 270, f. A, B (preocc. Papilio menentina Cramer, [1777])
Anapheis aurota ; [BRU, 160]
Belenois aurota ; [EBW]; [BOR, 146]; [AfrL]
Anaphaeis aurota ; [BMAT: 18, pl. 6, f. 27-29]
mesentina ; [BIR, 425]

Larva on Capparis oleoides, Maerua triphylla [PBSA], Capparis herbacea [BRU], Capparis spinosa, C. sepiaria, Capparis atphylla, C. heyneana, C. galeata, Maerua crassifolia, M. arenaria, Boscia senegalensis, B. albitrunca [BMAT]

India, Sikkim, Tropical Africa, Madagascar Belenois aurota aurota ; [BOR, 146]; [BAFR, 78]; [BK, 137]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 13, pl. 43, f. 1
?mesentina r. mesentina ; [BIR, 425]
Belenois (Belenois) aurota aurota ; [MadL]
B. a. augusta (Olivier, 1801)
Belenois aurota augusta ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 4
Ceylon Belenois aurota tabrobana ; [BOR, 146]
mesentina r. taprobana ; [BIR, 425]
Belenois aurota ?ssp. ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 14, pl. 43, f. 2-3 (Lmandraka, Madagascar)
B. a. turanica (Sheljuzhko, 1908)
S.Turkmenia, Tajikistan

Belenois creona (Cramer, [1776])
Tropical Africa (dry parts), Comoros, Madagascar, SW.Arabia. See [About maps] African Common White African Caper
Papilio creona Cramer, [1776]; Uitl. Kapellen 1 (8): 148, pl. 95, f. B, C
Belenois creona ; [EBW]; [AfrL]
Senegal - Nigeria, Sudan, W.Ethiopia Belenois creona creona ; [BAFR, 79 (text)]; [AfrL]
B. c. benadirensis (Storace, 1948)
Somalia Belenois creona benadirensis ; [BAFR, 79 (text)]; [AfrL]
B. c. boguensis (C. & R. Felder, 1865)
Agamsa?, Ethiopia Belenois creona boguensis ; [BAFR, 79 (text)]; [AfrL]
B. c. elisa (van Vollenhoven, 1869)
Comoro Is. Belenois creona elisa ; [BAFR, 79 (text)]; [AfrL]
B. c. leucogyne Butler, 1885
SW.Arabia Belenois creona leucogyne ; [BAFR, 79 (text)]; [AfrL]
B. c. prorsus (Talbot, 1943)
Madagascar Anapheis creona prorsus Talbot, 1943; Ann. Mag. nat. Hist. (11) 10: 305, TL: Madagascar, Maintirano
Belenois creona prorsus ; [BAFR, 79 (text)]; [AfrL]
Belenois creona prorsa ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 5
Belenois (Belenois) creona prorsus ; [MadL]
B. c. severina (Stoll, 1781)
South Africa - E.Africa, Zaire Papilio severina Stoll, 1781; in Cramer, Uitl. Kapellen 4 (29-31): 95
Belenois severina ; [EBW]
Belenois creona severina ; [BAFR, 79]; [BK, 137]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 7-9; [AfrL]

Larva on Capparis oleoides, C. zeyheri, Maerua triphylla [PBSA]

?B. c. infida Butler, 1888²

Belenois gidica (Godart, [1819])
Natal, Swaziland, Transvaal, Moçambique, Rhodesia, South West Africa, Tropical Africa (dry zones). See [About maps] African Veined White Pointed Caper
Pieris gidica Godart, [1819]; Encyclopédie Méthodique. 9 (1): 131
Belenois gidica ; [EBW]; [BAFR, 86]; [BK, 138]; [AfrL]

Larva on Capparis citrifolia [PBSA], Boscia, Maerua [BK]

Belenois gidica gidica ; [AfrL]
B. g. hypoxantha (Ungemach, 1932)
Belenois gidica hypoxantha ; [AfrL]
B. g. abyssinica (Lucas, 1852)
Belenois gidica abyssinica ; [AfrL]


22.7.2008 (29)

References:
[About maps]
Warning! The maps are automaticly generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); e-mail: Mark C. Williams
[BAFR]
D'Abrera, 1980; Butterflies of the Afrotropical Region
[BAUR]
D'Abrera, 1977; Butterflies of the Australian Region (2nd edn)
[BIR]
Wynter-Blyth, M. A., 1957; Butterflies of the Indian Region; (1982 Reprint)
[BK]
Larsen, Torben B., 1996; The Butterflies of Kenya and their natural history
[BMAT]
Tennent, John, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOR]
D'Abrera, B., 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III
[BOW]
Lewis, H. L., 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU]
Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997; Guide to the Butterflies of Russia and adjacent territories; Hesperiidae, Papilionidae, Pieridae, Satyridae; Volume 1
[Don Herbison-Evans]
and Stella A. Crossley; Lepidoptera Larvae of Australia; e-mail: Don Herbison-Evans; e-mail: Don Herbison-Evans
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[MadL]
Lees, D.C., Kremen, C. and Raharitsimba, H. 2003; in S.M. Goodman & J. P. Benstead (eds.), The Natural History of Madagascar; Classification, Diversity and Endemism of the Butterflies (Papilionoidea and Hesperioidea): A Revised Species Checklist: : pp. 762-793; University of Chicago Press. 1709 pp.
[PBSA]
Dickson, C.G.C (ed.), Kroon, D.M.; Pennington's Butterflies of Southern Africa; AD. DONKER 1978
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Boisduval, 1833
Faune Entomologique de Madagascar, Bourbon et Maurice. Lépidoptéres Faun. ent. Madagascar, Lépid.: 122pp, 16pls
Boisduval, 1836
Histoire Naturelle des insectes: Spécies général des lépidoptères Hist. nat. Ins., Spec. gén. Lépid. 1: 690pp, 24pls
Cramer, [1775]; Cramer, [1776]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 1 (1-7): 1-132, pl. 1-84 (1775), (8): 133-156, pl. 85-96 (1776)
Cramer, [1780]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 3 (17-21): 1-104, pl. 193-252 (1779), (22): 105-128, pl. 253-264 ([1780]), (23-24): 129-176, pl. 265-288 (1780)
Drury, [1773]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 2: 90pp, 50pls
K. L. & L. E. Dunn, 1991
Review of Australian Butterflies: distribution, life history and taxonomy. Pushlished by authors, Melbourne Review Austr. Butts. (1-4):
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Linnaeus, 1768
Sparrman (Thesis), Iter Chinense in Sparrman, Amoenit. acad., 7 (150): 497-506
Stoll, 1781
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Suffert, 1904
Neue afrikansche Tagfalter Dt. Ent. Z. Iris 17: 12-107, pl.
Trimen, 1883
Descriptions of twelve new species of South African Lepidoptera Rhopalocera Trans. ent. Soc. Lond. 1883: 347-363
Winhard, 2000
Pieridae I Butterflies of the world, 10: (1) 40pp, (2) 48pls


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxononic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.