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Colias Fabricius, 1807
Clouded Yellows
Colias Fabricius, 1807; Magazin f. Insektenk. (Illiger) 6: 284, TS: Papilio hyale Linnaeus
Eurymus Horsfield, [1829]; Descr. Cat. lep. Ins. Mus. East India Coy (2): 134, TS: Papilio hyale Linnaeus
= ; [NL4A, 102]
Ganura Zetterstedt, [1839]; (unavail.)
= ; [NL4A, 102]
Scalidoneura Butler, 1871; Proc. zool. Soc. Lond. 1871: 250, TS: Scalidoneura hermina Butler
= ; [NL4A, 102]
Eriocolias Watson, 1895; Entomologist 28: 167, TS: Papilio edusa Fabricius
= ; [NL4A, 102]
Coliastes Hemming, 1931; Entomologist 64: 273, TS: Papilio hyale Linnaeus
= ; [NL4A, 102]
Protocolias Petersen, 1963
= ; [NL4A, 102]
Mesocolias Petersen, 1963
= ; [NL4A, 102]
Neocolias Berger, 1986
= ; [NL4A, 102]
Palaeocolias Berger, 1986
= ; [NL4A, 102]
Eucolias Berger, 1986
= ; [NL4A, 102]
Similicolias Berger, 1986
= ; [NL4A, 102]
Paracolias Berger, 1986
= ; [NL4A, 102]
Colias, NSG Voucher Specimen [Wahlberg]
Colias, Pieridae Holarcticae [Heiner Ziegler]
 Colias ; Godman & Salvin, [1889], Biol. centr.-amer., Lep. Rhop. 2: 151; Winhard, 2000, Butterflies of the world, 10: 9; [LCG]; [NL4A, 102]; [AfrL]
fiColias hyale (Linnaeus, 1758)
S.Russia, Altai, Mongolia, NW.China. See [About maps] Vaaleakeltaperhonen Pale Clouded Yellow Ljusgul höfjäril Vaxfjäril Goldene Acht Le Soufre Szlaczkoń siarecznik
Papilio hyale Linnaeus, 1758; [GNB: 469], TL: S.England
Colias hyale, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias hyale, Svenska fjärilar [SVF]
Colias hyale, Moths and Butterflies of Europe [MBE]
Colias hyale, Extracts from [BRU] by [Ilya Osipov]
Colias hyale, Europaische Schmetterlinge [Mario Maier]
Colias hyale, Captain's European Butterfly Guide [Simon Coombes]
 Colias hyale ; [BRU, 172]; [EBW]; [BOW: pl. 4, f. 4-5]; [MRS, 220]; [Otakar Kudrna]
[hyale-1mv.jpg] 1024x768 (~234Kb) underside male Eiskeller, Berlin, Germany, 10-94, Photo © S. Shuichi Haupt [hyale-2fv.jpg] 462x400 (~33Kb) underside female The Irtysh River left bank floodland within the Omsk City, West Siberia, Russia. 27th August 1996 (on Cirsium setosum (Willd.) M.B.), Photo © Oleg Kosterin [hyale-3vU.jpg] identification uncertain 800x534 (~92Kb) underside Ukraine, Ivano Frankovsk, near Kalush, Stankova, 24.10.2001, Photo © Erich Mangl
[hyale-4vU.jpg] identification uncertain 1000x1162 (~159Kb) underside Russia, Udmurtia, Izhevsk (parklike areas along shore of "Izhevsky Prud"), 5.8.2004, Photo © Markku Savela [hyale-5fuv.jpg] 536x768 (~141Kb) upperside underside female Russia, Kaluga area, 09.09.2005, Photo © D. Smirnov [hyale-5muv.jpg] 586x768 (~141Kb) upperside underside male Russia, Kaluga area, 09.09.2005, Photo © D. Smirnov
[hyale-fv.jpg] 500x373 (~31Kb) underside female Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes [hyale.jpg] 600x448 (~67Kb) France, Pyrenees Orientales, Axat, Summer 1996, Photo © Simon Coombes

Larva on Vicia cracca [SPRK], Fabaceae, Vicia, Coronilla, Medicago, Lotus, Cytisus, Trifolium [BRU]


C. h. hyale
Europe, Ukraine, Kirghizia, Kazkhstan, S.Russia aegra Verity, 1947; Le Farfalle diurn. d'Italia 3: 262
= ; [LCG, 40]
alba Derenne, 1933; Lambillionea 33: 184
= ; [LCG, 40]
ab. albescens Metschl, 1922; Mitt. munchn. ent. Ges. 12: 6
= ; [LCG, 40]
albinotica Goodson, 1960; Ent. Gaz. 11: 18
= ; [LCG, 40]
alboradiata Herzig, 1935; Int. ent. Z. 29: 365
= ; [LCG, 40]
antehyale Verity, 1947; Le Farfalle diurn. d'Italia 3: 264
= ; [LCG, 40]
anteubercalida Verity, 1947; Le Farfalle diurn. d'Italia 3: 266
= ; [LCG, 40]
apicata Tutt, 1896; Brit. Butts: 253
= ; [LCG, 40]
appendiculata Verity, 1947; Le Farfalle diurn. d'Italia 3: 260
= ; [LCG, 40]
argentea Fritsch, 1913; Ent. Rundsch. 30: 46
= ; [LCG, 40]
argyphea Lowe, 1909; Ent. Rec.: 36
= ; [LCG, 40]
atava Reutti, 1853; Ubers. Lep. Fauna Baden: ?
= ; [LCG, 40]
bipupillata Cabeau, 1912; Rev. mens. Soc. ent. Namur 12: 54
= ; [LCG, 40]
brabantica Strand, 1912; : 49-50
= ; [LCG, 40]
brevis de Crombrugghe, 1911; Rev. mens. Soc. ent. Namur 11: 104
= ; [LCG, 40]
canarina Stauder, 1922; Mitt. münch. ent. Ges. 12 (1-6): 25
= ; [LCG, 40]
cleopatra Latier, 1935; P.-v. Soc. linn. Bordeaux 86: 149
= ; [LCG, 40]
crassipuncta Le Charles, 1927; Bull. Soc. ent. Fr. 8: 138
= ; [LCG, 41]
crocoexpansa Caruel, 1955; Revue Française de Lépid. 1-2: 27
= ; [LCG, 41]
deannulata Rocci, 1920; Atti Soc. Lig. 30 (4): 30
= ; [LCG, 41]
demarginata Nitsche, 1913; Verh. zool.-bot. Ges 63: 21
= ; [LCG, 41]
deuara Herzig, 1935; Int. ent. Z. 29: 365
= ; [LCG, 41]
duplex Ksienschopolski, 1911; Trudy obsch. Izsl. Volyni Zitomir 8: 28, pl. 1, f. 2
= ; [LCG, 41]
elongata Vorbrodt & Müller-Rutz, 1917; Mitt. schweiz. ent. Ges. 12 (9-10): 434
= ; [LCG, 41]
emarginata Röber, [1907]; in Seitz, Grossschmett. Erde 1: 65
= ; [LCG, 41]
eratoides Strand, 1918; Soc. ent. 33 (7): 27, TL: Turkestan
= ; [LCG, 41]
flava Husz, 1880; Jb. Ungar Karpath. Ver. 8: 272
= ; [LCG, 41]
flavida Gruber, 1932; Ent. Zeit. 46: 195, f. 14
= ; [LCG, 41]
ab. flavoapicalis Metschl, 1922; Mitt. munchn. ent. Ges. 12: 7
= ; [LCG, 41]
flavofasciata Lambillion, 1907; Cat. Lep. Belg. (2nd ed): 409
= ; [LCG, 41]
flavopupillata Soja, 1929; Ver. zool.-bot. Ges. Wien. 79: 107
= ; [LCG, 41]
ab. flavoradiata Metschl, 1922; Mitt. munchn. ent. Ges. 12: 7
= ; [LCG, 41]
gartneri Skala, 1912; Ver. naturf. Ver. Brünn 50: 63
= ; [LCG, 41]
gavalgnii Stauder, 1921; Z. Wiss. InsekBriol. 16: 220
= ; [LCG, 41]
gloriciana Fritsch, 1918; Ent. Rundsch. 35: 6
= ; [LCG, 41]
grisandrocontata Lempke, 1954; Tijdschr. Ent. 97: 311
= ; [LCG, 41]
griseoapicata Herzig, 1935; Int. ent. Z. 29: 365
= ; [LCG, 41]
heliceides de Selys de Longchamps, 1844; Mém. Soc. roy. Sci. Liége 2: 30
= ; [LCG, 41]
immaculata Derenne, 1925; Rev. Mens. Soc. ent. Namur 25: 92
= ; [LCG, 41]
impunctata Rocci, 1920; Atti Soc. Lig. 30 (4): 30
= ; [LCG, 41]
infrastriata Lempke, 1954; Tijdschr. Ent. 97: 306
= ; [LCG, 41]
inornatura Derenne, 1923; Rev. Mens. Soc. ent. Namur 23: 39
= ; [LCG, 41]
intermedia Tutt, 1896; Brit. Butts: 253
= ; [LCG, 41]
inversa Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 434
= ; [LCG, 41]
junior Geest, 1905; Z. wiss. InsekBiol. 1: 380
= ; [LCG, 41]
macropuncta Finke, 1934; Int. ent. Z. 28: 293, TL: Germany, Göttingen
= ; [LCG, 41]
magnoimaculata Lempke, 1954; Tijdschr. Ent. 97: 314
= ; [LCG, 41]
melanina Verity, [1909]; Rhopalocera Palaearctica 1: 223
= ; [LCG, 41]
melaina Mück, 1919; Ent. Zeit. 33 (9): 35, TL: Bâle
= ; [LCG, 42]
mellaertsi Lambillion, 1913; Rev. Mesn. Soc. Ent. Namur 13: 126
= ; [LCG, 42]
metschli Gruber, 1932; Ent. Zeit. 46: 195, f. 15
= ; [LCG, 42]
minor van Mellaerts, 1926; Lambillionea 26: 84
= ; [LCG, 42]
myrmhyale Meeke, 1926; Int. ent. Z.: 173
= ; [LCG, 42]
nigriformis Finke, 1934; Int. ent. Z. 28: 394, TL: Germany
= ; [LCG, 42]
nigripuncta Caruel, 1955; Revue Française de Lépid. 1-2: 27
= ; [LCG, 42]
nigrofasciata Grum-Grshimailo, 1884; in Romanoff, Mém. Lép. 1: 163-164, TL: Saratov
= ; [LCG, 42]
nigrosparsata Lempke, 1954; Tijdschr. Ent. 97: 312
= ; [LCG, 42]
obsoleta Tutt, 1896; Brit. Butts: 259
= ; [LCG, 42]
ocellata Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83: 70
= ; [LCG, 42]
ochrocretacea Fritsch, 1918; Ent. Rundsch. 35: 6
= ; [LCG, 42]
omnimarginata Hafner, 1912; Carniola: 221, pl. 3, f. 41
= ; [LCG, 42]
opposita Zusanek, 1918; Z. öst. EntVer. 3: 36
= ; [LCG, 42]
ab. pallescens Metschl, 1922; Mitt. munchn. ent. Ges. 12: 6
= ; [LCG, 42]
pallida Robson & Gardner, 1886; Young Naturalist, suppl.: 4
= ; [LCG, 42]
pallida Cockerell, 1889; Entomologist 22: 4 [?]
= ; [LCG, 42]
pallidior Cockerell, 1912; Entomologist, 45: 323
= ; [LCG, 42]
parisiensis Oberthür, 1936; Lambillionea 36: 90
= ; [LCG, 42]
parvipuncta Lempke, 1954; Tijdschr. Ent. 97: 313
= ; [LCG, 42]
posticomunita Lempke, 1954; Tijdschr. Ent. 97: 307
= ; [LCG, 42]
postrema Rocci, 1920; Atti Soc. Lig. 30 (4): 29
= ; [LCG, 42]
ab. pseudohelice Metschl, 1922; Mitt. munchn. ent. Ges. 12: 6
= ; [LCG, 42]
pupillata Lempke, 1936; Lambillionea, 36: 103
= ; [LCG, 42]
pygmaea Lambillion, 1906; Rev. Mens. Soc. ent. namur. 6: 22
= ; [LCG, 42]
minor Vorbrodt, 1912; in Vorbrodt & Müller-Rutz, Die Schmett. Scweitz 1: 30
= ; [LCG, 42]
radiata Geest, 1905; Z. wiss. InsekBiol. 1: 379
= ; [LCG, 42]
radiiformis Schultz, 1904; Nyt. Mag. naturvid. 42: 41
= ; [LCG, 42]
rufa Verity, [1909]; Rhopalocera Palaearctica 1: 222, pl. 41, f. 1
= ; [LCG, 42]
sagittata Schnaider, 1950; Polsk. Pismo. Ent. 19: 240
= ; [LCG, 42]
schonfeldi Mezger, 1926; Ent. Anz. 6: 18
= ; [LCG, 42]
sieversoides Verity, [1909]; Rhopalocera Palaearctica 1: 221, pl. 40, f. 34
= ; [LCG, 42]
simplex Neuburger, 1905; Soc. ent. 20 (6): 42
= ; [LCG, 42]
striata Zusanek, 1918; Z. öst. EntVer. 3: 36
= ; [LCG, 42]
striata Romaniszyn, 1926; Polsk. Pismo ent. 5: 144
= ; [LCG, 42]
submarginata Finke, 1934; Int. ent. Z. 28: 394, TL: Germany
= ; [LCG, 42]
subobsoleta Pionneau, 1936; Echange 52: 23
= ; [LCG, 42]
sufflava Verity, 1947; Le Farfalle diurn. d'Italia 3: 262
= ; [LCG, 43]
supervacanea Krulikovsky, 1908; Soc. Ent. 23 (1): 3
= ; [LCG, 43]
trimaculata Steiner, 1935; Int. etn. Z. 29: 269
= ; [LCG, 43]
uber Verity, 1926; Ent. Rec. 38: 171
= ; [LCG, 43]
ubercalida Verity, 1947; Le Farfalle diurn. d'Italia 3: 266
= ; [LCG, 43]
uhli Kovats, 1899; Ent. Zeit. 12: 169
= ; [LCG, 43]
unimaculata Tutt, 1896; Brit. Butts: 253
= ; [LCG, 43]
venata Lempke, 1936; Lambillionea, 36: 76
= ; [LCG, 43]
vernalis Verity, [1909]; Rhopalocera Palaearctica 1: 222, pl. 47, f. 32
= ; [LCG, 43]
viridescens Sloper, 1903; Wheeler. Bull. Switz.: 69
= ; [LCG, 43]
viridis van Mellaerts, 1926; Lambillionea 26: 84
= ; [LCG, 43]
Colias hyale hyale ; Winhard, 2000, Butterflies of the world, 10: 9, pl. 11, f. 1-2
Colias poliographus inversa ;
Colias hyale hyale ; [LCG]
C. h. altaica Verity, 1911
Altai, Sayan hyale altaica Verity, 1911; Rhopalocera Palaearctica 1: 348, pl. 69, f. 4-5
Colias hyale altaica ; [BRU, 172]
Colias hyale hyale f. altaica ; [LCG, 40]
C. h. alta Staudinger, 1886
Tian-Shan, Ghissar, Darvaz, Alai, Pamirs hyale alta Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 200, TL: Kirgizia, Alai Mts.
Colias poliographus alta ; [BRU, 173]
Colias hyale alta ; [LCG, 43]
C. h. palidis Fruhstorfer, 1910
East Siberia f. palidis Fruhstorfer, 1910
palides[sic] ; [LCG, 44 (missp.?)]
Colias hyale palidis ; [BRU, 172]
C. h. irkutskana Stauder, 1924
Transbaikalian hyale irkutskana Stauder, 1924; Mitt. münch. ent. Ges. 14: 62, TL: Irkutsk, Ojek
f. fulvocoma Krulikovsky, 1907; Revue Russe Ent. 7: 29, TL: Viatka
= ; [CG]
Colias hyale irkutskana ; [BRU, 172]; [LCG, 43]
C. h. novasinensis Reissinger, 1989
Gansu Colias hyale novasinensis Reissinger, 1989; Neue Ent. Nachricten 26: 199, TL: Gansu, Xining
Colias hyale novasinensis ; [LCG]
Colias sareptensis Staudinger, 1871
SEU, CEU, S.Russia, Asia Minor, Caucasus, Transcaucasia - Middle Asia - China. See [About maps] Berger's Clouded Yellow Sydlig ljusgul höfjäril Szlaczkoń południowiec
Colias sareptensis Staudinger, 1871; in Staudinger & Wocke, Cat. Lepid. europ. Faunegeb.: 5, TL: Sarepta
Colias alfacariensis Ribbe, 1905; Soc. ent. 20 (18): 137, TL: Andalusia
Colias australis Verity, 1911; Rhopalocera Palaearctica 1: 347, TL: Andalusia, Spain
calida Verity, 1916; Entomologist's Rec. J. Var. 28: 99
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis ubercalida Reissinger, 1972; : 112
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis senonica Reissinger, 1972; Atalanta 3: 359
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis paracalida Reissinger, 1972; Atalanta 3: 366
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis orthocalida Reissinger, 1974; Atalanta 5: (1-33)
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis kantaraica Reissinger, 1989; Neue Ent. Nachricten 26: 223
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis bergeri Reissinger, 1989; Neue Ent. Nachricten 26: 126
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis antebergeri Reissinger, 1989; Neue Ent. Nachricten 26: 126
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis postbergeri Reissinger, 1989; Neue Ent. Nachricten 26: 126
= Colias alfacariensis ; [LCG, 45]
anteorthocalida Reissinger, 1989; Neue Ent. Nachricten 26: 82
= Colias alfacariensis ; [LCG, 48]
postorthocalida Reissinger, 1898; : 82
= Colias alfacariensis ; [LCG, 48]
Colias alfacariensis rumilica Reissinger, 1989; Neue Ent. Nachricten 26: 129
= Colias alfacariensis ; [LCG, 48]
Colias alfacariensis slavonica Reissinger, 1989; Neue Ent. Nachricten 26: 154
= Colias alfacariensis ; [LCG, 50]
Colias alfacariensis magyarica Reissinger, 1989; Neue Ent. Nachricten 26: 162
= Colias alfacariensis ; [LCG, 46]
Colias alfacariensis metacalida Reissinger, 1989; Neue Ent. Nachricten 26: 170
= Colias alfacariensis ; [LCG, 47]
Colias alfacariensis hyrcanica Reissinger, 1989; Neue Ent. Nachricten 26: 133
= Colias alfacariensis ; [LCG, 46]
Colias alfacariensis mandarina Reissinger, 1989; Neue Ent. Nachricten 26: 189
= Colias alfacariensis ; [LCG, 47]
Colias alfacariensis, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias alfacariensis, Moths and Butterflies of Europe [MBE]
Colias sareptensis, Extracts from [BRU] by [Ilya Osipov]
Colias australis, Europaische Schmetterlinge [Mario Maier]
Colias alfacariensis, Captain's European Butterfly Guide [Simon Coombes]
 Colias australis ; [EBW]
Colias alfacariensis ; [BMAT: 101, pl. 32, f. 14-16]; Winhard, 2000, Butterflies of the world, 10: 9, pl. 11, f. 3-6; [LCG, 44]; [Otakar Kudrna]
Colias sareptensis ; [BRU, 172]

Larva on Coronilla varia, Hippocrepis comosa, Astragalus monspessulanus [LCG, (for alfacariensis)]

C. s. sareptensis
Colias alfacariensis ; [LCG, 49 (form)]
C. s. vihorlatensis Reissinger, 1989
Carpathians Colias alfacariensis vihorlatensis Reissinger, 1989; Neue Ent. Nachricten 26: 166
= Colias alfacariensis ; [LCG, 50]
C. s. remota Reissinger, 1989
SEU, Caucaus Colias alfacariensis remota Reissinger, 1989; Neue Ent. Nachricten 26: 149
= Colias alfacariensis ; [LCG, 48]
Colias sareptensis remota ; [BRU, 172]
C. s. fontainei Reissinger, 1989
Armenian Highland, Talysh, Kopet-Dagh Colias alfacariensis fontainei Reissinger, 1989; Neue Ent. Nachricten 26: 141
= Colias alfacariensis ; [LCG, 45]
C. s. saissanica Reissinger, 1989
SE.Kazakhstan Colias alfacariensis saissanica Reissinger, 1989
= Colias alfacariensis ; [LCG, 49]
Colias sareptensis saissanica ; [BRU, 172]
Colias erate (Esper, [1805])
-. See [About maps] Eastern Pale Clouded Yellow Szlaczkoń erate
Papilio erate Esper, [1805]; Ausl. Schmett. 1 (2): ?13, pl. 119, f. 3, TL: Sarepta, S. Russia
Colias erate, Moths and Butterflies of Europe [MBE]
Colias erate, Extracts from [BRU] by [Ilya Osipov]
 Colias erate ; [BRU, 173]; [EBW]; [BOW: pl. 4, f. 1-2]; [MRS, 218]; [AfrL]; [Otakar Kudrna]
[erate-1mv.jpg] 951x624 (~56Kb) underside male Sukayu, Aomori, Japan, 8-90, Photo © S. Shuichi Haupt [erate-2v.jpg] 800x707 (~67Kb) underside Japan: Lake Kawaguchiko, Yamanashi Prefecture, 23.08.2007, Photo © Peter Payzant

Larva on Medicago, Trifolium, Onobrychis, Melilotus, (etc.) [BRU]


C. e. erate
Ukraine, Turkey, Lebanon, Bulgaria, Romania, Macedonia, Greece, Hungary, Austria, Turkmenistan, Kazakhstan, Kirhizia, Uzbekistan, Tadzhikistan, Afghanistan Colias neriene Fischer von Waldheim, 1823; Entomographia Imp. ross., 2: 251, pl. 11, f. 3-4
afghana O. Bang-Haas, 1927; Horae Macrolep. Palaearct. 1: 42, TL: Afghanistan, Paghman Mts.
= ; [LCG, 52]
albida Rühl, [1893]; Palaearktischen Grosschmetterlinge: 157
= ; [LCG, 52]
androconiata Jachontov, 1910; Revue Russe Ent. 9 (4): 380
= ; [LCG, 52]
beckeri Gerhard, 1882; Berl. ent. Z. 26: 125
= ; [LCG, 52]
benesignata Sheljuzhko, 1913; Dt. Ent. Z. Iris 27: 20
= ; [LCG, 52]
chryseis Röber, [1907]; in Seitz, Grossschmett. Erde 1: 66, pl. 26b
= ; [LCG, 52]
chrysodona Boisduval, 1840; Genera Index Eur. Lepid.: 7
= ; [LCG, 52]; [LCG, 52]
chrysohyaleoides Verity, [1908]; Rhopalocera Palaearctica 1: 219
chrysopallida Verity, [1908]; Rhopalocera Palaearctica 1: 221, pl. 57, f. 29
= ; [LCG, 52]
chrysothemoides Verity, [1908]; Rhopalocera Palaearctica 1: 219, pl. 47, f. 25
= ; [LCG, 52]
citronea Mezger, 1932; Lambillionea, 32: 4
= ; [LCG, 52]
conjuncta Verity, [1908]; Rhopalocera Palaearctica 1: 219
= ; [LCG, 52]
costaenigrata Mezger, 1932; Lambillionea, 32: 4
= ; [LCG, 52]
diffusa Verity, [1908]; Rhopalocera Palaearctica 1: 219, pl. 40, f. 30
= ; [LCG, 52]
flavomaculata Mezger, 1932; Lambillionea, 32: 4
= ; [LCG, 52]
garica Verity, 1911; Rhopalocera Palaearctica 1: 348, pl. 69, f. 6-7
= ; [LCG, 52]
helichta Lederer, 1853; Verh. zool.-bot. Ver. Wien 2: 33 (spell.?)
= ; [LCG, 52 (as helichtha[sic])]
helichta Talbot, 1935; Lep. Cat. 66: 423
= Colias erate erate f. androconiata Jachontov, 1910; [LCG, 52]
hyaleoides Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4: 321, TL: Fergana
= ; [LCG, 52]
maculigena Avinoff, 1910; Horae Soc. Ent. Ross. 39: 229, TL: Borokoro
= ; [LCG, 52]
pallida Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 3
= ; [LCG, 52]
parva Rühl, [1893]; Palaearktischen Grosschmetterlinge: 157
= ; [LCG, 52]
semifaillaei Pionneau, 1928; Echange 44 (431): 3
= ; [LCG, 52]
unimaculata Pionneau, 1928; Echange 44 (431): 3
= ; [LCG, 52]
Colias erate erate f. chrysodona ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 10
Colias erate erate f. chrysopallida ; Winhard, 2000, Butterflies of the world, 10: 9, pl. 11, f. 9
Colias erate erate f. hyaleoides ; Winhard, 2000, Butterflies of the world, 10: 9, pl. 11, f. 8
Colias erate erate ; Winhard, 2000, Butterflies of the world, 10: 9, pl. 11, f. 7; [LCG, 51]
C. e. amdensis Verity, 1911
China (Qinghai, Gansu, Sichuan) erate amdensis Verity, 1911; Rhopalocera Palaearctica 1: 348, pl. 41, f. 8, TL: Amndo, Kukunor
chrysocoma Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27: 379, TL: Sichuan
= ; [LCG, 52]
hera Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27: 379, TL: Sichuan
= ; [LCG, 52]
Colias erate amdensis ; [LCG, 52]
C. e. marnoana Rogenhofer, 1884
Sudan, Ethiopia, SW.Arabia Colias erate marnoana Rogenhofer, 1884; Verh. zool.-bot. Ges. Wien 33: 22, TL: Sudan
androides Storace, 1956; Ann. Mus. civ. Stor. nat. Genova 68: 349
= ; [LCG, 54]
solilucis Berger, 1953; Entomologist 86: 172
= ; [LCG, 54]
Colias erate marnoana ; [BAFR, 54]; [LCG, 53]; [AfrL]
C. e. sinensis Verity, 1911
Yunnan?, Mandschuria, N.Korea Colias erate sinensis Verity, 1911; Rhopalocera Palaearctica 1: 349, pl. 69, f. 19-22, TL: Kioi-Kiang, Chang-Yang
Colias erate sinensis ; [MRS, 218]; [LCG, 55]
C. e. formosana Shirôzu, 1955
Taiwan Colias erate formosana Shirôzu, 1955; Sieboldia 1 (3): 229, pl. 30, f. 1-4
Colias erate formosana ; [MRS, 218]; [LCG, 52]

Larva on Trifolium pratense, T. repens, Medicago polymorpha, M. sativa, Lotus australis, Astragalus sinicus [MRS]

C. e. lativitta Moore, 1882
Nepal, N.India erate lativitta Moore, 1882; Proc. zool. Soc. Lond. 1882 (1): 255, TL: Nepal
erate gigantea Verity, 1911; Rhopalocera Palaearctica 1: 347, pl. 72, f. 25 (preocc. Colias gigantea Strecker)
Colias erate tania Hemming, 1933; Entomologist 66: 278 (repl. erate gigantea Verity, 1911)
= ; [LCG, 53]
glicia Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 164 (2 April, 1910)
= ; [LCG, 53]
kachgarica Verity, 1911; Rhopalocera Palaearctica 1: 348, pl. 69, f. 6-7
= ; [LCG, 53]
Colias erate lativitta ; [LCG, 53]
C. e. poliographus Motschulsky, 1860
Japan, Amur, Ussuri, Sakhalin, Tian Shan Colias poliographus Motschulsky, 1860; Etud. Ent., 9: 29, TL: "Khokodody", Japan
Colias simoda de l'Orza, 1869; Lep. Jap.: 16
= ; [LCG, 55]
immamis Verity, 1911; Rhopalocera Palaearctica 1: 349, pl. 41, f. 10
= ; [LCG, 55]
murina Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 164 (2 April, 1910)
= ; [LCG, 55]
napata Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 164 (2 April, 1910)
= ; [LCG, 55]
pyxagathus Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 164 (2 April, 1910)
= ; [LCG, 55]
Colias hyale zuzuki Matsumura, 1908; Ent. mon. Mag. 2: 120, pl. 5, f. 8, TL: Kyoto
= ; [LCG, 55]
Colias poliographus, Extracts from [BRU] by [Ilya Osipov]
 Colias poliographus ; [BRU, 173]
Colias erate poliographus ; Huang, 2001; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 11-12; [LCG, 54]

Larva on Trifolium, Vicia, (etc.) Glycine max [BRU], Parochetus communis, Lucerne spp. [MRS]

C. e. tomarias Bryk, 1942
Kuriles Colias hyale tomarias Bryk, 1942; Dt. Ent. Zeit. Iris, 56: 12, TL: Kurile Is., Kumashir & Tomari
tokotana Bryk, 1942; Dt. Ent. Zeit. Iris, 56: 12
Colias poliographus tomarias ; [BRU]
Colias erate tomarias ; [LCG, 55]
C. e. naukratis Fruhstorfer, 1909
Altai, S.Siberia, Transbaikalia erate naukratis Fruhstorfer, 1909; in Seitz, Gross-Schmett. Erde 9 : 164 (2 April, 1910)
Colias poliographus naukratis ; [BRU]
Colias erate naukratis ; [LCG, 54]
C. e. nilagiriensis C. & R. Felder, 1859
S.India erate nilagiriensis C. & R. Felder, 1859; Wien. ent. Monats. 3 (12): 395, TL: S.India, Nilgiri Mts.
Colias erate nilagiriensis ; [LCG, 54]
Colias libanotica Lederer, 1858
-. See [About maps] Colias libanotica Lederer, 1858; Wien. ent. Monats. 2 (5): 140, pl. 2, f. 1-2, TL: Syria [Lebanon]
C. l. libanotica
Israel, Lebanon, Turkey hybrida Verity, [1909]; Rhopalocera Palaearctica 1: 259
= ; [LCG, 56]
offusa Verity, [1909]; Rhopalocera Palaearctica 1: 259
= ; [LCG, 56]
pulchior Verity, [1909]; Rhopalocera Palaearctica 1: 259
Colias libanotica libanotica ; [LCG, 56]
C. l. heldreichi Staudinger, 1862
Greece Colias libanotica heldreichi Staudinger, 1862; Stett. Ent. Ztg. 23: 257, TL: Mt. Veluchi, Greece
fountainei Aigner-Abafi, 1901; Rovart. Lap. 8: 30, TL: Mt. Chelmos
= Colias libanotica heldreichii ; [LCG, 57]
griseoviridis Verity, [1909]; Rhopalocera Palaearctica 1: 260-274, pl. 45, f. 29, TL: Greece
= Colias libanotica heldreichii ; [LCG, 57]
Colias aurorina heldreichi ;
Colias libanotica heldreichii f. fountainei ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 14
Colias libanotica heldreichii ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 13; [LCG, 56]
C. l. sovarensis Blom, 1979
Iran Colias aurorina sovarensis Blom, 1979; Entomol. Ber. Amst. 39 (1): (10-14), TL: Iran, 8km N-E Khosh-Yeilaq, Kuh-e-Sovar Mts, 2000-2500m
Colias libanotica sovarensis ; [LCG, 57]
Colias aurorina Herrich-Schäffer, 1850
-. See [About maps] Dawn Clouded Yellow
Colias aurorina Herrich-Schäffer, 1850; Syst. Bearb. Schmett. Europ. 1: f. 453-456, TL: Armenia
Colias aurorina, Moths and Butterflies of Europe [MBE]
Colias aurorina, Extracts from [BRU] by [Ilya Osipov]
 Colias aurorina ; [BRU, 177]; [Otakar Kudrna]

Larva on Astragalus [H&R], Astracantha caucasica, A. denudata [BRU, Nekrutenko, 1990]

C. a. aurorina
Armenia, Georgia, Azerbaijan, Talysh, N.Iran, Turkey alba Rühl, [1893]; Palaearktischen Grosschmetterlinge: 731
= ; [LCG, 58]
aurantiaca Rühl, [1893]; Palaearktischen Grosschmetterlinge: 730
= ; [LCG, 58]
deserticola Verity, [1909]; Rhopalocera Palaearctica 1: 258
= ; [LCG, 58]
Colias aurorina auronina ; [LCG, 57]
C. a. daghestanica Verhulst, 1994
Daghestan, Armenia Colias aurorina daghestanica Verhulst, 1994; Lambillionea 94: 115, TL: Daghestan, Rutul
Colias aurorina daghestanica ; [LCG, 58]
C. a. kermana Eckweiler, 1979
Iran (Kerman) Colias aurorina kermana Eckweiler, 1979; Ent. Zs. Frankf. a. M. 89 (3): 29-32, TL: Iran, Kerman, Jiroft-Rayen, 2800-3000m
Colias auronina kermana ; [LCG, 59]

Larva on Astragalus sp. [LCG, 59]

C. a. rosei Gross & Ebert, 1975
N.Iran (Elburs) Colias aurorina rosei Gross & Ebert, 1975; J. ent. Soc. Iran, suppl. 1: 11, TL: Iran, Elburs, Damavand
Colias aurorina rosei ; [LCG, 59]
C. a. taurica Rebel, 1901
Turkey Colias aurorina f. taurica Rebel, 1901; Verh. zool.-bot. Ges. Wien, 51: 310, TL: Turkey, Taurus
leucothea Rebel, 1917; Sber. Akad. Wiss. Wien (126) 1: 251
= ; [LCG, 60]
Colias aurorina taurica ; [LCG, 59]
C. a. transcaspica Christoph, 1889
Turkmenistan (Kopet-Dagh, W.Transcaspia) aurorina transcaspica Christoph, 1889; in Romanoff, Mém. Lépid. 5: 5, pl. 1, f. 1, TL: W.Transcaspia, Nuchur
chrysocoma Eversmann, 1851; Bull. Soc. imp. Nat. Moscou 24 (2): 622, TL: Caucasia
= ; [LCG, 60]
tamara Nordman, 1851; : 413, TL: Caucasia
= ; [LCG, 60]
Colias aurorina transcaspica ; [LCG, 60]
Colias sagartia Lederer, 1869
N.Iran. See [About maps] Colias sagartia Lederer, 1869; Hor. Soc. ent. ross. 6 (2): 75, TL: Iran
daubii Geest, 1905; Z. wiss. InsekBiol. 1: (378-381)
= ; [LCG, 61]
lisa Geest, 1905; Z. wiss. InsekBiol. 1: (378-381)
= ; [LCG, 61]
pullata Neuburger, 1905; Soc. ent. 20 (6): 42
= ; [LCG, 61]
Colias sagartia ; [LCG, 60]

Larva on Acantholimon, Astragalus spp. [LCG, 61]

Colias chlorocoma Christoph, 1888
Transcaucasia, Turkey, Iran. See [About maps] Colias chlorocoma Christoph, 1888; Horae Soc. ent. Ross. 22 (3-4): 308, TL: Turkey
Colias chlorocoma, Extracts from [BRU] by [Ilya Osipov]
 Colias chlorocoma ; [BRU, 177]

Larva on Onobrychis cornuta [BRU, Nekrutenko, 1990]

C. c. chlorocoma
Turkey ochracea Verhulst, 1993; Lambillionea 93 (1): 44
= ; [LCG, 62]
sulphurea Miller, 1912; Dt. ent. Z. Iris 26 (4): 223
= ; [LCG, 62]
Colias chlorocoma chlorocoma ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 15; [LCG, 61]

Larva on Astragalus aurens, A. lycioides [LCG, 62]

C. c. aladagensis Verhulst, 1993
Turkey Colias chlorocoma aladagensis Verhulst, 1993; Lambillionea 93 (1): 44, TL: Turkey, Nidge, Aladglari, Pinarbasi 200-2600m
Colias chlorocoma aladagensis ; [LCG, 62]

Larva on Astragalus angusitfolius? [LCG, 63]

C. c. tkatschukovi O. Bang-Haas, 1936
Armenia, Azerbaijan chlorocoma tkatschukovi O. Bang-Haas, 1936; Ent. Z. Frankf. a. M. 50: 287, TL: Armenia, Daralagaes 2000m, 45.30E, 39.25N
Colias chlorocoma tkatschukovi ; [LCG, 63]
C. c. wyatti Häuser & Schurian, 1978
Iran chlorocoma wyatti Häuser & Schurian, 1978; Atalanta, 9 (3): (240-244), TL: Iran, Dugijan
Colias chlorocoma wyatti ; [LCG, 63]
Colias caucasica Staudinger, 1871
W.Caucasus, NE.Turkey. See [About maps] Balkan Clouded Yellow
Colias caucasica Staudinger, 1871; in Staudinger & Wocke, Cat. Lepid. europ. Faunegeb.: 6, TL: vicinity of Akhaltsikhe, Georgia
Colias caucasica, Extracts from [BRU] by [Ilya Osipov]
 Colias caucasica ; [BRU, 178]

Larva on Cytisus [BRU, Nekrutenko, 1990]

C. c. caucasica
anna Gerhard, 1882; Berl. ent. Z. 26: 125
= ; [LCG, 65]
olga Romanoff, 1882; Horae Soc. ent. ross. 17: 127, pl. 4, f. 1-3
= ; [LCG, 65]
alba Rühl, [1893]; Palaearktischen Grosschmetterlinge: 162
= ; [LCG, 65 (= olga)]
Colias caucasica caucasica ; [LCG, 64]

Larva on Astragalus caucasicus [LCG, 65]

Colias balcanica Rebel, 1901
Bosnia, Yugoslavia, Bulgaria, Greece. See [About maps] Colias Myrmidone v. Balcanica Rebel, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 1: 250, TL: Bosnia and Herzegovina, Vucja Bara
anna Schawerda, 1913; Verh. zool.-bot. Ges. Wien 63: 144
= ; [LCG, 66]
aphrodite Schawerda, 1908; Jber. Wien. ent. Ver. 19: 89
= ; [LCG, 66]
caltha Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71: 148
= ; [LCG, 66]
daphnis Schawerda, 1913; Verh. zool.-bot. Ges. Wien 63: 144
= ; [LCG, 66]
isabellina Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71: 149
= ; [LCG, 66]
immaculata Schawerda, 1922; : 148
= ; [LCG, 66]
leucorhorion Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71: 148
= ; [LCG, 66]
magnomaculata Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71: 148
= ; [LCG, 66]
micans Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71: 148
= ; [LCG, 66]
nicholli Schawerda, 1907; Verh. zool. bot. Ges. Wien 57: 219
= ; [LCG, 66]
nigrocuneata Schawerda, 1908; Jber. Wien. ent. Ver. 19: 89
= ; [LCG, 66]
nigrovittata Rebel, 1922; Verh. zool.-bot. Ges. Wien 71: 31
= ; [LCG, 67]
pseudobalcanica Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 77
= ; [LCG, 67]
rebeli Schawerda, 1906; Verh. zool. bot. Ges. Wien 56: 651
= ; [LCG, 67]
rosea Schawerda, 1919; Z. öst. EntVer. 4 (7): 67
= ; [LCG, 67]
scotina Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71: 148
= ; [LCG, 67]
semialba Wagner, 1907; Ent. Zeit. 21 (20): 125
= ; [LCG, 67]
balcanica ; Abadjiev, 2002, Neue Ent. Nachr. 23: 16
Colias caucasica balcanica ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 16; [LCG, 65]
Colias balcanica ; [Otakar Kudrna]

Larva on Chamaecytisus hirsutus, Chamaecytisus absinthioides [LCG, 66]

Colias myrmidone (Esper, 1781)
-. See [About maps] Danube Clouded Yellow Orangeroter Heufalter Szlaczkoń szafraniec
Papilio myrmidone Esper, 1781; Die Europ. Schmett. 1 (2): pl. 65, f. 1-2, TL: Turnau, Hungary
Colias myrmidone, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias myrmidone, Extracts from [BRU] by [Ilya Osipov]
Colias myrmidone, Europaische Schmetterlinge [Mario Maier]
 Colias myrmidone ; [BRU, 178]; [Otakar Kudrna]

Larva on Cytisus [H&R]

C. m. myrmidone
Germany, Poland, Ukraine, Romania, Hungary, Slovakia, Austria, Bulgaria, Caucasus, SW.Russia aegra Verity, [1909]; Rhopalocera Palaearctica 1: 265
= ; [LCG, 67]
agnes Pieszczek, 1905; Verh. zool.-bot. Ges. Wien: 416
= ; [LCG, 67]
alba Staudinger, 1871; in Staudinger & Wocke, Cat. Lepid. europ. Faunegeb.: 6
= ; [LCG, 67]
amicans Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 73, pl. 1, f. 2
= ; [LCG, 67]
angustimarginata Metschl, 1925; Int. Ent. Zeit. 19: 25
= ; [LCG, 67]
antihygiaea Mayer, 1916; Ent. Zeit. 29 (26): 102
= ; [LCG, 67]
aurantiaca Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 75, pl. 2, f. 18
= ; [LCG, 67]
bahri Skala, 1912; Int. ent. Z. 6 (3): 16
= ; [LCG, 68]
bicolor Metschl, 1922; Mitt. munchn. ent. Ges. 12: 12
= ; [LCG, 68]
caliginosa Klos, 1914; Mitt. naturw. Ver. Steiermark 50: 214
= ; [LCG, 68]
callos Maurer, 1911; Jber. Wien ent. Ver. 22: 217
= ; [LCG, 68]
chrysothemides Verity, 1911; Rhopalocera Palaearctica 1: 358, pl. 72, f. 1-2
= ; [LCG, 68]
completa Pionneau, 1937; Echange 53: 10
= ; #
demarginata Metschl, 1922; Mitt. munchn. ent. Ges. 12: 8
= ; [LCG, 68 (= opposita)]
discolor Fritsch, 1912; Ent. Rundsch. 29 (21): 143
= ; [LCG, 68]
edusaeformis Klemensiewicz, 1910; Spraw. Kom fizyogr. 45: 50
= ; [LCG, 68]
edusoides Skala, 1912; Int. ent. Z. 6 (3): 16
= ; [LCG, 68]
pallida Zelezny, 1916; Z. öst. ent. Ver. 1: 20
= Colias myrmidone myrmidone f. edusoides ; [LCG, 68]
flavescens Garbowski, 1892; Sber. Akad. Wiss Wien.: 903
= ; [LCG, 68]
griseomarginata Berger, 1911; Jahrb. Wien. ent. Ver. 21: 77
= ; [LCG, 68]
depunctata Nitsche, 1912; Verh. zool.-bot. Ges. Wien 62: 109
= ; [LCG, 68 (= hartmanni Jouki, 1907)]
hartmanni Jouki, 1907; Cas. csl. Spol. ent. 4: 77
= ; [LCG, 68]
helma Geest, 1905; Z. wiss. InsekBiol. 1: 379
= ; [LCG, 68]
impunctata Verity, [1909]; Rhopalocera Palaearctica 1: 264
= ; [LCG, 68]
inspersa Metschl, 1922; Mitt. munchn. ent. Ges. 12: 11
= ; [LCG, 68]
intermedia Maurer, 1905; Verh. zool.-bot. Ges. Wien: 148
= ; [LCG, 68]
inumbrata Schultz, 1904; Nyt. Mag. naturvid. 42: 43, TL: Austria
= ; [LCG, 68]
lutea Zelezny, 1915; Ent. Rdsch. 32: 59
= ; [LCG, 58]
mediocarnea Metschl, 1922; Mitt. munchn. ent. Ges. 12: 11
= ; [LCG, 58]
micans Röber, [1907]; in Seitz, Grossschmett. Erde 1: 69
= ; [LCG, 68]
nana Mayer, 1910; Int. ent. Z. 4: 182, TL: Graz
= ; [LCG, 68]
myrmidonides Ksienschopolski, 1911; Trudy obsch. Izsl. Volyni Zitomir 8: 30
= ; [LCG, 68 (nana Mayer, 1910)]
nigerrima Pieszczek, 1905; Verh. zool.-bot. Ges. Wien: 420
= ; [LCG, 68]
nigrovenata Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 75, pl. 2, f. 13, TL: Graz
= ; [LCG, 68]
obscura Skala, 1912; Ver. naturf. Ver. Brünn 50: 63
= ; [LCG, 68]
opposita Metschl, 1922; Mitt. munchn. ent. Ges. 12: 8
= ; [LCG, 68]
orcus Ronnicke, 1917; Z. Öst. EntVer. 2 (9): 77, pl. 3, f. 28
orcusides Strand, 1928; Arch. Naturgesch. 92 A (8): 51 (repl. orcus Ronnicke, 1917)
pallescens Fritsch, 1912; Ent. Rundsch. 29 (21): 143
= ; [LCG, 68]
pallida Pieszczek, 1912; Jber. Wien Ent. Ver. 22: 223
= ; [LCG, 68]
pallidomaculata Metschl, 1922; Mitt. munchn. ent. Ges. 12: 8
= ; [LCG, 68]
permaculata Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 76, pl. 2, f. 20
= ; [LCG, 68]
pieszezeki Predota, 1909; Jber. Wien ent. Ver. 20: 141
= ; [LCG, 68]
pseudomaureri Metschl, 1925; Int. Ent. Zeit. 19: 25
= ; [LCG, 68]
pseudorebeli Klos, 1906; : 651
= ; [LCG, 68]
pupillata Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 71
= ; [LCG, 68]
rima Metschl, 1922; Mitt. munchn. ent. Ges. 12: 11
= ; [LCG, 68]
rubroflammea Zelezny, 1916; Z. öst. ent. Ver. 1: 20
= ; [LCG, 68]
rubroisabellina Fritsch, 1912; Ent. Rundsch. 29 (21): 142
= ; [LCG, 68]
rudolfi Ronnicke, 1927; Z. Öst. EntVer. 12 (8): 76
= ; [LCG, 69]
schwabi Pieszcek, 1912; : 232
= ; [LCG, 69]
striata Pieszcek, 1917; : 71
= ; [LCG, 69]
striata Metschl, 1922; Mitt. munchn. ent. Ges. 12: 8 (preocc. striata Pieszcek, 1917)
unipuncta Pieszcek, 1922; : 8
= ; [LCG, 69]
verna Verity, 1911; Rhopalocera Palaearctica 1: 358
= ; [LCG, 69]
virescalis Stöckl, 1929; Polsk. Pismo ent. 7: 65
= ; [LCG, 69]
Colias myrmidone myrmidone ; [LCG, 67]; [LCG, 68]

Larva on Cytisus ratisbonensis, C. capitatus, Chamaecytisus ruthenicus [LCG, 67 (Gorbunov & Korshunov, 1995)]

C. m. ermak Grum-Grshimailo, 1890
S.Urals, SW.Siberia, NW.Kazakhstan myrmidone ermak Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4: 301, TL: SE.Ural
irinae Krulikovsky, 1909; Rev. russ. ent. 9: 110
= ; [LCG, 69]
sophiae Krulikovsky, 1909; Rev. russ. ent. 9: 110
= ; [LCG, 69]
stellata Krulikovsky, 1908; Soc. Ent. 23 (1): 3
= ; [LCG, 69]
vernalis Ksienschopolski, 1911; Trudy obsch. Izsl. Volyni Zitomir 8: 30
= ; [LCG, 69]
Colias myrmidone ermak ; [LCG, 69]
Colias nastes Boisduval, 1832
Altai, Sayan - Chukot Peninsula, Alaska, Greenland, Labrador, Greenland. See [About maps] Labrador Sulphur
Colias nastes Boisduval, 1832; Icon. hist. Lépid. Europe 1 (23-24): 245 (1834), (3-4): pl. 8 (1832), f. 4-5
Colias nastes, Lepidoptera collection [SZMN]
Colias nastes, Butterflies and Moths of North America [BAMONA]
Colias nastes, Norges Sommerfugler [NORLEP]
Colias nastes, Extracts from [BRU] by [Ilya Osipov]
 Eurymus nastes ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 11
Colias nastes ; [NACL, #4216]; [BRU, 170]; [Opler]
?Colias nastes ; [EBW, (or = tyche?)]
Colias nastes ; [BOW: pl. 4, f. 6 (or = tyche?)]

Larva on Astragalus alpinus, Oxytropis, Hedysarium [BRU, Scott, 1986]

C. n. nastes
Canada (Labrador, Northwest Territories, Baffin Island, Southampton Island) gueneei Avinoff, 1935; in Holland & Avinoff, Mem. Carn. Mus. 12 (2): 13
= ; [LCG, 70]
Colias nastes rossii Guenée, 1864; Ann. Soc. Ent. Fr. (4) 4: 199
= ; [LCG, 70]
Eurymus nastes rossii ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 11
Colias nastes rossii ; [NACL, #4216a]
Colias nastes nastes ; [LCG, 69]

Larva on Astragalus alpinum [LCG, 70]

C. n. aliaska O. Bang-Haas, 1927
Alaska, Canada (Yukon, Northwest Territories, Victoria Island, Banks Island) Colias nastes aliaska O. Bang-Haas, 1927; Horae Macrolep. Palaearct. 1: 41, pl. 5, f. 24-25, TL: Alaska, Ramport
subarctica McDunnough, 1928; Can. ent. 60: 270, TL: Bernard Harbour, N. W. T.
= ; [LCG, 71]
alaskae ; McDunnough, 1938, Mem. South. Calif. Acad. Sci. 1: 8 (missp.)
Colias nastes aliaska ; [NACL, #4216e]; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 17-18; [LCG, 70]
C. n. dezhnevi Korshunov, 1996
NE.Siberia (Magadan, Chukotka, Bilibino) Colias nastes dezhnevi Korshunov, 1996; Dopolneniya i ispravleniya k knige "Dnevnye babochki aziatskoi chasti Rossii": 18, TL: Siberia, Verchoiansk, Adytch River
Colias nastes dezhnevi, Lepidoptera collection [SZMN]
 Colias tyche dezhnevi ; Grieshuber, 1998
Colias nastes dezhnevi ; [LCG, 71]
C. n. dioni Verhulst, 1999
Canada (Alberta) Colias nastes dioni Verhulst, 1999; Lambillionea 99 (4): 624, TL: Canada, Alberta, Kananaskis Co., Plateau Mtn 2500m
Colias nastes dioni ; [LCG, 71]
C. n. jakutica Kurentzov, 1970
Far East (Yakutia) Colias nastes jacutica Kurentzov, 1970; Butterflies of the USSR Far East: ? (spell.?)
?Colias nastes jakutica Kurentzov, 1970; (spell.?)
Colias jacuticola Weiss & Mracek, 1989; Nachr. ent. Ver. Apollo 10 (4): 281 (spell.?)
= ; [LCG, 72]
?Colias nastes jakuticola Weiss & Mracek, 1989; (spell. ?)
Colias nastes jacutica, Lepidoptera collection [SZMN]
 Colias nastes jakutica ; [LCG, 72]
C. n. moina Strecker, 1880
Canada (Northwest Territories, Manitoba) moina Strecker, 1880; Bull. Brooklyn ent. Soc. 3 (5): 34, TL: Canada, N.Manitoba, Churchill
harperi (Gunder, 1932); Can. Ent. 64 (2): 278, TL: Ft. Churchill, Manitoba
Colias nastes moina ab. harperi ; [LCG, 73]
Eurymus moina ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 11
Colias nastes moina ; [NACL, #4216b]; [LCG, 73]
C. n. streckeri Grum-Grshimailo, 1895
Canada (Alberta, British Columbia), N.Washinginton, Montana streckeri Grum-Grshimailo, 1895; Horae Soc. ent. Ross. 29: 290, TL: Alberta, Laggan, alke Louise
sibirica Kurentzov, 1970; (nom. praeocc.)
obscurata Verity, 1911; Rhopalocera Palaearctica 1: 355, pl. 61, f. 6, TL: Alberta, Lake Louise
= ; [LCG, 74 (ab.)]
palliflava McDunnough, 1927; Canadian Entomologist, 59: 154
= ; [LCG, 74 (ab.)]
Eurymus nastes streckeri ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 11
Colias nastes streckeri ; [NACL, #4216c]; [LCG, 74]
C. n. zemblica Verity, 1911
Novaja Zemlja nastes zemblica Verity, 1911; REF#x: 355, pl. 71, f. 10, TL: Siberia, Novaja Zemlja
?zembalica Verity, 1911; (spell.?)
Colias tyche zemblica ; Grieshuber, 1998
Colias nastes zemblica ; [LCG, 74]
C. n. cocandicides Verity, 1911
Colias nastes cocandicides ; [NACL, #4216d]
C. n. ferrisi Verhulst, 2004
Alaska Colias nastes ferrisi Verhulst, 2004; Lambillionea 104 (1): 129, TL: Alaska
fiColias tyche (Böber, 1812)
. See [About maps] Tunturikeltaperhonen Pale Arctic Clouded Yellow Fjällfjäril
tyche (Böber, 1812); Mém. Soc. Imp. Nat. Moscou 3: 21, ?72, pl. 3, f. 3-6 [??ref, see melinos]
Colias melinos Eversmann, 1847; Bull. Soc. Imp. Nat. Moscou 20 (?2): 72, pl. 3, f. 3-6 [??ref, see tyche }
Colias tyche, Butterflies and Moths of North America [BAMONA]
Colias tyche, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias tyche, Svenska fjärilar [SVF]
Colias tyche, Moths and Butterflies of Europe [MBE]
Colias tyche, Extracts from [BRU] by [Ilya Osipov]
 Colias tyche ; [BRU, 170]; [Opler]
[tyche-1fv.jpg] 533x400 (~46Kb) underside female Open pine forest on a hill just NE of the Chita City, Transbaikalia, Siberia, Russia. 17th June 1995 (ssp. deckerti Verity, 1909), Photo © Oleg Kosterin

Larva on Astragalus frigidus [SPRK, given as C. nastes]

Larva on Astragalus alpinus, Oxytropis, Caragana [BRU]


C. t. tyche
N.Siberia, SE.Siberia, Altai, Sayan, Yakutia, Transbaikalia, Baikal, Amur, Mongolia deckerti Verity, [1909]; Rhopalocera Palaearctica 1: 236, pl. 42, f. 27-28, TL: SE.Siberia, Mt. Apfel
= ; [LCG, 75]
herzi Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 3: 16, TL: Viluy
= ; [LCG, 76]
jeholensis Matsumura, 1939; Bull. biogeogr. Soc. Japan 9 (20): 348, TL: Mongolia, Re He, Jehol, Kingan Mtn
= ; [LCG, 76]
ludmilla Hemming, 1933; Entomologist 66: 278 (repl. chryseis Verity, 1911
= ; [LCG, 76]
chryseis Verity, 1911; (praeocc. Colias erate chryseis Röber, 1907)
vitimensis Austaut, 1899; Le Naturaliste 21: 284
= ; [LCG, 76]
magna Rühl, [1893]; Palaearktischen Grosschmetterlinge: 153, TL: ?Lake Baikal
= ; [LCG, 75]
melinoides Verity, 1926; Ent. Rec. 38: 171
= ; [LCG, 75]
montana Verity, 1911; Rhopalocera Palaearctica 1: 354, pl. 70, f. 34-38
= ; [LCG, 75]
viridis O. Bang-Haas, 1915; Dt. ent. Z. Iris 29: 99, TL: Mts Nan Shan, China
= ; [LCG, 75]
Colias tyche montana ;
Colias tyche tyche ; [LCG, 74]
C. t. relicta Kurentzov, 1970
Chukot Peninsula, Far East Colias melinos relicta Kurentzov, 1970; Butterflies of the USSR Far East: ?, TL: NE.Siberia, R. Okhota
Colias tyche relicta ; [LCG, 77]
fiC. t. werdandi Zetterstedt, 1839
NEU tyche werdandi Zetterstedt, 1839; Insecta Lapponica : 908, TL: Torne-Lappland
alba Schulte, 1953; Ent. Zeit. 62: 150
= ; [LCG, 78]
anastigma Röber, [1907]; in Seitz, Grossschmett. Erde 1: 63
= ; [LCG, 78]
bruneomaculata Schulte, 1953; Ent. Zeit. 62: 151
= ; [LCG, 78]
castaneapunctata Sheldon, 1912; Entomologist 45: 27
= ; [LCG, 78]
christiernssoni Lampa, 1885; Ent. Tidskr. 6: 11
= ; [LCG, 78]
connexa Sheldon, 1912; Entomologist 45: 27
= ; [LCG, 78]
ellingeri Schulte, 1953; Ent. Zeit. 62: 151
= ; [LCG, 78]
flavopunctata Sheldon, 1912; Entomologist 45: 27
= ; [LCG, 78]
grisea Schulte, 1953; Ent. Zeit. 62: 151
= ; [LCG, 78]
immaculata Lampa, 1885; Ent. Tidskr. 6: 11
= ; [LCG, 78]
insignata Thurau, 1903; Berl. ent. Z. 48: 113
= ; [LCG, 78]
mariannae Schulte, 1953; Ent. Zeit. 62: 152
= ; [LCG, 78]
nigrita Schulte, 1953; Ent. Zeit. 62: 152
= ; [LCG, 78]
palaeonides Schulte, 1953; Ent. Zeit. 62: 151
= ; [LCG, 78]
phicomonides Verity, 1911; Rhopalocera Palaearctica 1: 355, TL: Norway
= ; [LCG, 78]
radiata Thurau, 1903; Berl. ent. Z. 48: 113
= ; [LCG, 78]
sulphurea Lampa, 1885; Ent. Tidskr. 6: 10
= ; [LCG, 78]
torneoensis Sheldon, 1912; Entomologist 45: 27
= ; [LCG], TL: 78;#
Colias tyche werdandi ; [LCG, 77]
Colias werdandi ; [Otakar Kudrna]

Larva on Astragalus alpinus, Vaccinium myrtillus, Oxytropis lappona [LCG, 77]

C. t. magadanica Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001
Magadan Colias tyche magadanica Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001; Helios, Coll. lep. articles II: (103-115)
C. t. flinti Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001
Wrangel Is. Colias tyche flinti Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001; Helios, Coll. lep. articles II: (103-115)
C. t. kolosovae Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001
E.Chukotka Colias tyche kolosovae Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001; Atalanta 32 (1/2): 262 (repl. Colias tyche olga Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001)
Colias tyche olga Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001; Helios, Coll. lep. articles II: (103-115) (preocc. Colias olga Romanoff, 1882)
Colias phicomone (Esper, 1777)
-. See [About maps] Mountain Clouded Yellow Grünlicher Heufalter
Papilio phicomone Esper, 1777; Die Europ. Schmett. 1 (2): pl. 56, f. 1-2, TL: Styria
Colias phicomone, Moths and Butterflies of Europe [MBE]
Colias phicomone, Europaische Schmetterlinge [Mario Maier]
Colias phicomone, Captain's European Butterfly Guide [Simon Coombes]
 Colias phicomone ; [EBW]; [BOW: pl. 4, f. 8]; [Otakar Kudrna]
C. p. phicomone
Alps, N.Carpathians, N.Italy alpiumnitida Verity, 1926; Ent. Rec. 38: 171
= ; [LCG, 79]
anticopupillata Verity, 1947; Le Farfalle diurn. d'Italia 3: 269
= ; [LCG, 79]
bipupillata Mezger, 1932; Lambillionea, 32: 210, TL: Simplon
= ; [LCG, 79]
blachieri Culot, 1909; Bull. Soc. Lép. Genève 1: 265
= ; [LCG, 79]
derosea Wagner, 1926; Z. Öst. Ent.-Ver. Wien 11: 25
= ; [LCG, 79]
elegans Schultz, 1904; Nyt. Mag. naturvid. 42: 42
= ; [LCG, 79]
fasciata Ronnicke, 1920; Ent. Zeit. 34: 70
= Colias phicomene phicomene ; [LCG], TL: 79;#
flavida Osthelder, 1913; Mitt. münchn. ent. Ges. 4: 28
= ; [LCG, 79]
flava Osthelder, 1925; Mitt. münchn. ent. Ges. 15 (suppl.): 67
= ; [LCG, 78 (= flavida Osthelder, 1913)]
Colias phicomone f. fleischmanni Stauder, 1924; Int. ent. Z. 18: 37
= ; [LCG, 79]
geesti Neuburger, 1905; Soc. ent. 20 (6): 42
= ; [LCG, 79]
gynomorpha Verity, 1911; Rhopalocera Palaearctica 1: 353, pl. 70, f. 22
= ; [LCG, 79]
albidior Ronnicke, 1920; Ent. Zeit. 34: 71
= ; [LCG, 79 (= gynomorpha Verity, 1911)]
hyalides Verity, 1911; Rhopalocera Palaearctica 1: 353, pl. 70, f. 24
= ; [LCG, 79]
incompleta Mezger, 1934; Lambillionea, 34: 54, TL: Switzerland, Valais, Rieffelalp
= ; [LCG, 79 (ab.)]
lanceolata Verity, 1947; Le Farfalle diurn. d'Italia 3: 270, TL: Dolomites, Alpi Agordine, Falcade, Ponte de la Schita 2000m
= ; [LCG, 79]
obscura Ronnicke, 1920; Ent. Zeit. 34: 71
= ; [LCG, 79]
pallida Hoffmann & Klos, 1914; Mitt. nat. Ver. Steierm. 50: 208
= ; [LCG, 79]
passa Verity, 1911; Rhopalocera Palaearctica 1: 353, pl. 70, f. 23
= ; [LCG, 79]
periphaes Fruhstorfer, 1908; Ent. Zs. 22 (12): 51
= ; [LCG, 79]
phaedra Schawerda, 1924; Mitt. münchn. ent. Ges. 14: 79
= ; [LCG, 79]
porrecta Heinrich, 1930; Ent. Zeit. 43: 295, TL: Zürich
= ; [LCG, 79]
privata Ronnicke, 1920; Ent. Zeit. 34: 71
= ; [LCG, 79]
pulverulenta Verity, 1926; Ent. Rec. 38: 171, TL: Cottian Alps
= ; [LCG, 79]
pupillata Refhous, 1909; Bull. Soc. Lép. Genève 1: 26, pl. 8
= ; [LCG, 79]
pupillata Ronnicke, 1920; Ent. Zeit. 34: 71
= ; [LCG, 79 (= pupillata Refhous, 1909)]
unimaculata Pionneau, 1924; Echange 39: 23
= ; [LCG, 79 (= pupillata Refhous, 1909)]
saturata Austaut, 1903; Ent. Zeit. 18: 143
= ; [LCG, 79]
theia Schawerda, 1919; Z. öst. EntVer. 4 (7): 67
= ; [LCG, 79]

Larva on Vicia sativa, Vicia pirenaica, V. angustifolia, V. narbonensis, Medicago sativa, Coronilla sp., Lotus corniculatus, Oxytropis montana, Hippocrepis comosa [LCG, 78]

C. p. juliana Hospital, 1948
Cantabria phicomone juliana Hospital, 1948; Trab. Mus. Cienc. nat. Barcelona 1 (2): 11, TL: Cordillera Cantabrica, Picos de Europa
impunctata Verity, [1909]; Rhopalocera Palaearctica 1: 230, pl. 62, f. 3, TL: Asturites, Picos de Europa, 2000m
= ; [LCG, 80]
Colias phicomone juliana ; [LCG, 79]
C. p. oberthueri Verity, [1909]
Pyrenees phicomone oberthueri Verity, [1909]; Rhopalocera Palaearctica 1: 231, pl. 42, f. 5-6, TL: Pyrenees, Vernet-les-Bains, Gèdre
Colias phicomone oberthueri ; [LCG, 80]
fiColias palaeno (Linnaeus, 1761)
. See [About maps] Suokeltaperhonen Moorland Clouded Yellow Palaeno Sulphur Arctic Sulfur Svavelgul höfjäril Svavelfjäril Hochmoorgelbling Szlaczkoń torfowiec
Papilio palaeno Linnaeus, 1761; Fauna Suecica (Edn 2): 272, TL: Uppsala, Sweden
Colias palaeno, Lepidoptera collection [SZMN]
Colias palaeno, Butterflies and Moths of North America [BAMONA]
Colias palaeno, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias palaeno, Svenska fjärilar [SVF]
Colias palaeno, Norges Sommerfugler [NORLEP]
Colias palaeno, Moths and Butterflies of Europe [MBE]
Colias palaeno, Extracts from [BRU] by [Ilya Osipov]
Colias palaeno, Europaische Schmetterlinge [Mario Maier]
 Eurymus palaeno ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 10
Colias palaeno ; [NACL, #4218]; [BRU, 179]; [EBW]; [BOW: pl. 4, f. 7]; [MRS, 219]; [Otakar Kudrna]
[palaeno-1vf.jpg] 600x645 (~34Kb) underside female FINLAND: Li: Utsjoki Ailigas, 775:50, 14.7.1998, Photo © Markku Savela [palaeno-2.jpg] 750x499 (~34Kb) Sipoo 670:40, 18.06.1980, Photo © Pertti Pakkanen

Larva on Vaccinium uliginosum [SPRK] [MRS], Vaccinium uliginosum, V. caespitosum [NAB]


C. p. palaeno
Finland, Sweden, Norway, Estonia lapponica Staudinger, 1871; in Staudinger & Wocke, Cat. Lepid. europ. Faunegeb.: 5
philomene Hübner, 1803; Samml. eur. Schmett. [1] : f. 602-603 (1805)
avinoffi Verity, 1911; Rhopalocera Palaearctica 1: 346, TL: Finland
= ; [LCG, 81]
cretacea Schilde, 1884; Ent. Nachr. 10: (335-346)
= ; [LCG, 81]
nordströmi Bryk, 1923; Ent. Tidskr. 44: 109
= ; [LCG, 81]
octava Bryk, 1923; Ent. Tidskr. 44: 108
= ; [LCG, 81]
parva Hoyningen-Huene, 1901; Sber. Naturf. Ges. Dorpat (Jurjew) 12: 460, TL: Estonia
= ; [LCG, 81]
schildei Staudinger, 1892; Dt. Ent. Z. Iris, 5: 310
= ; [LCG, 81]
Colias palaeno palaeno ; [LCG, 81]
Misidentified/misapplied:
?Eurymus palaeno werdandi Herrich-Schäffer, 1848; Dyar, 1903, Bull. U.S. natn. Mus., 52: 10
werdandi Herrich-Schäffer, 1848; Schmett. Eur. 1: 41-42, f. 403-404

Larva on Vaccinium uliginosum, V. myrtillus [LCG, 81]

C. p. aias Fruhstorfer, 1903
Japan palaeno aias Fruhstorfer, 1903; Dt. ent. Z. Iris 16 (1): 48, TL: Japan
sugitanii Esaki, 1929; Zephyrus 1 (2): 42, f. 3-4
= ; [LCG, 82]
nekkana Matsumura, 1939; Bull. biogeogr. Soc. Japan 9 (20): 348
= ; [LCG, 82]
Colias palaeno aias ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 19; [LCG, 81]
C. p. europome (Esper, 1778)
Germany, Belgium, Slovakia, Romania, Ukraine palaeno europome (Esper, 1778); Die Schmett. 1 (1): pl. 42, f. 1-2, TL: Saxony
atavista Stephan, 1923; Dt. Ent. Z. Iris 37: 30
= ; [LCG, 82]
aurantiaca Hackray, 1934; Lambillionea 34: 193
= ; [LCG