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Pyronia Hübner, [1819]
Pyronia Hübner, [1819]; Verz. bek. Schmett. (4): 59, TS: Pyronia tithone Hübner
Idata de Lesse, 1952; Ann. Soc. ent. Fr. 121: 72, TS: Epinephele ida var. cecilia Vallentin
Pasiphana de Lesse, 1952; Ann. Soc. ent. Fr. 121: 72, TS: Papilio bathseba Fabricius
Pyronia [Wahlberg]
Pyronia (Maniolina) ; [Wahlberg]

Pyronia tithonus (Linnaeus, 1771)
CEU, SEU, Asia Minor, Caucasus, Morocco (Ketama). See [About maps] Gatekeeper Hedge Brown l'Amaryllis Gelbe Ochsenauge Lobito agreste Przestrojnik titonus Buskgräsfjäril Rödgul Slåttergräsfjäril
Papilio tithonus Linnaeus, 1771; Mantissa Plant. 2: 537
Pyronia tithone Hübner, [1819]; Verz. bek. Schmett. (4): 59
Epinephele tithonus distincta Rothschild, 1933; Novit. Zool. 38 : 321, TL: Ketama (Morocco)
Pyronia tithonus [Kimmo Silvonen]
Pyronia tithonus [Mario Maier]
Pyronia tithonus [Simon Coombes]
Pyronia tithonus ; [EBW]; [BOW: pl. 8, f. 12, 17]; [BMAT: 82, pl. 28, f. 1-4]
Maniola tithonus ; [Otakar Kudrna]
[tithonus-1fu.jpg] 329x500 (~37Kb) upperside female Bois de Caurel, Brittany, France 11/07/95, Photo © Simon Coombes [tithonus-1mu.jpg] 297x450 (~57Kb) upperside male Wembury, Devon, UK 03/07/90, Photo © Simon Coombes

Larva on Gramineae, Poa annua, Milium [H&R], (in Spain) Dactylis glomerata, Elytrigia repens, Milium effusum, Brachypodium sp., Lolium sp. [BMAT]


distincta
[distincta-1u.jpg] 600x435 (~33Kb) upperside Morocco, Djebel Tisirène, Rif, Photo © Michel Tarrier



Pyronia cecilia (Vallantin, 1894)
Morocco, SEU, Asia Minor. See [About maps] Southern Gatekeeper
Epinephele ida var. cecilia Vallantin, 1894; Le Naturaliste 16: 260, TL: Algeria
Papilio ida Esper, 1784; Die Schmett. Th. I, Bd. 2 (9): 184, pl. 92, f. 2
Papilio ida Esper, 1794; Die Schmett., Suppl. Th 1 (3-4): 52 (1789), (5-6): pl. 102 (1794), f. 3
Epinephele ida lapidepta Seitz, 1908; [BMAT: 138, pl. 46, f. [d]], TL: S.Algeria [El Kantara]
Pyronia cecilia [Wahlberg]
Pyronia cecilia [Simon Coombes]
Epinephele ida latipicta ; Oberthür, 1925, Etud. Lépid. Comp., 23: 85 (missp. lapidepta Seitz)
Idata cecilia ;
Pyronia cecilia ; [BMAT: 83, pl. 28, f. 19-22]; [BOW: pl. 8, f. 3]
Maniola cecilia ; [Otakar Kudrna]
[cecilia-1v.jpg] 600x435 (~27Kb) underside Morocco, Pays Beni-Routen, Rif, Photo © Michel Tarrier

Larva on Poaceae, Deschampsia caespitosa [BMAT]



Pyronia bathseba (Fabricius, 1793)
Morocco, Algeria, SWEU. See [About maps] Spanish Gatekeeper Lobito listado
Papilio bathseba Fabricius, 1793; Ent. Syst. 3 (1): 235, TL: Barbarie (N.Africa)
Papilio pasiphae Esper, 1781; Die Schmett. Th. I, Bd. 2 (3): 99, pl. 67, f. 4 (preocc. Papilio pasiphae Cramer, [1775])
Epinephele pasiphae var. philippina Austaut, 1877; Petites Nouv. Ent. 2 (176): 149, TL: Nemours [Ghazaouet] (Algeria)
Epinephele pasiphae var. taurina Oberthür, 1915; Étud. Lépid. Comp. 10 : 359, pl. 288, f. 2335, TL: Magenta (Algeria)
Epinephele pasiphae var. multiocellata Oberthür, 1915; Étud. Lépid. Comp. 10 : 359, pl. 288, f. 2337, TL: Djurdjura (Algeria)
Pyronia bathseba pardilloi de Sagarra, 1924; Bull. Inst. catal. Hist. nat. (2) 4: 199, TL: Barcelona
Pyronia bathseba [Wahlberg]
Pyronia bathseba [Mario Maier]
Pyronia bathseba [Simon Coombes]
Pyronia bathseba ; [BOW: pl. 8, f. 9]; [BMAT: 82, pl. 28, f. 5-18]
Maniola bathseba ; [Otakar Kudrna]
[bathseba-1u.jpg] 500x408 (~33Kb) upperside Mt. Canigou, Pyrenees Orientales, France 27/7/96, Photo © Simon Coombes [bathseba-1v.jpg] 500x388 (~38Kb) underside Mt. Canigou, Pyrenees Orientales, France 27/7/96, Photo © Simon Coombes [bathseba-2v.jpg] 600x435 (~51Kb) underside Morocco, Chefchaouen, Rif, Photo © Jean Delacre
[bathseba-3v.jpg] 600x435 (~43Kb) underside Morocco, Cirque de Jaffar, Haut Atlas, Photo © Michel Tarrier

Larva on Gramineae, Brachypodium [H&R]



Pyronia coenonympha Felder, 1865²
Himalayas. See [About maps] coenonympha Felder, 1865²; Reise der ... Fregate Novara: 492

Pyronia janiroides (Herrich-Schäffer, [1851])
Algeria, Tunisia. See [About maps] False Meadow Brown
Epinephele janiroides Herrich-Schäffer, [1851]; Syst. Bearb. Schmett. Europ. 1 (51): (ii) pl. 111, f. 533
Epinephele janiroides Herrich-Schäffer, [1851]; Syst. Bearb. Schmett. Europ. 6 (48): 17
Epinephele janiroides ab. abbreviata Seitz, 1906; Gross-Schmett. Erde 1 : 138, pl. 46, f. b, TL: Algeria
?[Satyrus] janiroides [?f.] tisamene Boisduval
[Satyrus] janiroides [?f.] tisamene ; Oberthür, 1876, Étud. d'Ent. 1 : 28
Pyronia janiroides ; [BOW: pl. 8, f. 11]; [BMAT: 83, pl. 28, f. 23-27]

Larva on Poa annua [BMAT]



25.7.2009 (5)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BMAT]
Tennent, John, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOW]
Lewis, H. L., 1974; Butterflies of the World; ISBN 0 245 52097 X
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[H&R]
Higgins, L.G, Riley, N.D, 1970; A Field Guide to the Butterflies of Britain and Europe
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Jean Delacre]
http://homepage.mac.com/jdelacre
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Esper, 1781; Esper, 1784
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Fortsetzung. Band 2 Die Schmett. Th. I, Bd. 2 (1): 1-36, pl. 51-56 (1780) (2): 37-68, pl. 57-62 (1781) (3): 69-100, pl. 63-68 (1781) (4): 101-124, pl. 69-74 (1781) (5): 125-140, pl. 75-80 (1782) (7): 141-156, pl. 81-82 (1783) (8): 157-172, pl. 83-88 (1783) (9): 173-184, pl. 89-92 (1784) (28): 185-190, pl. 93 (1786)
Esper, 1794
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 1. Abschnitt 1 Die Schmett., Suppl. Th 1 (1-2): 1-8, pl. 94 (1789) (3-4): 9-60, pl. 95-101 (1789) (5-6): 61-68, pl. 102-106 (1794) ([7]): [69-88], pl. [107-109] (1798) (8-9): 89-104, pl. 110-112 (1800) (10): 105-120, pl. 113-116 ([1803-1804])
Herrich-Schäffer, [1851]
Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu J. Hübner's Sammlung europäischer Schmetterlinge, 1843 (-1855), Die Tagfalter Syst. Bearb. Schmett. Europ. 1 Text (ii) Plates _Papilionides_ (iii) Plates _Hesperides_ (9): Title ("1843") (1): [1-3], 4-16, (ii) pl. 1-5, f. 1-29 (1843) (2): 17-40, (ii) pl. 29-34, f. 131-156 (1843) (3): 41-56, (ii) pl. 6-12, f. 30-60, pl. 35-36, f. 157-164 (1844) (4): 57-80, (ii) pl. 13-22, f. 61-107 (1844) (5): 81-90, (ii) pl. 23-28, f. 108-130, pl. 37-41, f. 165-189 (1844) (6): 91-98, (ii) pl. 42-44, f. 190-203 (1844) (7): 99-114, (ii) pl. 45-52, f. 204-248 (1844) (9): 115-122 (1844) (10): 123-146, (ii) pl. 53-57, f. 249-274 (1845) (11): 147-162 (1845) (12): (iii) pl. 1-3, f. 1-17 (1845) (13): 163-164, (ii) pl. 58-65, f. 275-315 (1846) (14): (ii) pl. 66-70, f. 316-338, (iii) pl. 4-5, f. 18-30 (1846) (26): (ii) pl. 71-77, f. 339-376 (1847) (32): (ii) pl. 78-82, f. 377-400 (1848) (36): (ii) pl. 83-89, f. 401-426, (iii) pl. 6, f. 31-36 (1848) (37): (ii) pl. 90-91, f. 427-437 (1849) (43): (ii) pl. 92-94, f. 438-450, 452-453 (1850) (45): (ii) pl. 95-102, f. 453-489 (1850) (47): (ii) pl. 103, f. 490-495 (1851) (49): (ii) pl. 104-109, f. 496-528 (1851) (51): (ii) pl. 110-118, f. 529-570 (1851) (54): (ii) pl. 119-123, f. 571-596 (1852) (56): (ii) pl. 124-127, f. 597-618, pl. 130-132, f. 628-639, pl. 134, f. 646-651 (1852) (59): (ii) pl. 128-129, f. 619-627, pl. 133, f. 640-645 (1853) (65): (iii) pl. 7, f. 37-42, (index) [1], 2-20 (1854) (66): (index) 21-24 (1855)
Herrich-Schäffer, [1851]
Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu J. Hübner's Sammlung europäischer Schmetterlinge, Sechter un letzter Band, 1843-1856 Syst. Bearb. Schmett. Europ. 6 (i) Erkl. Taf. Umrissen [Macrolep.] (ii) Erläuterung Taf. Umrissen Microlep. Nachtrag zum ersten Bande (iv) Syst. Lep. Europae (1): (i) pl. [I] (1843) (3): (i) [I]-II, pl. II-IV (1844) (6): (i) pl. V (1844) (7): (i) pl. VI (1844) (8): (i) III-X, pl. VII-VIII (1844) (9): (i) pl. IX-XI (1844) (11): (i) pl. XII (1845) (13): (i) XI-XIV, pl. XIII-XIV (1846) (17): (i) pl. XVI (1846) (22): (ii) pl. I-III (1847) (35): (i) pl. XV (1848) (36): (i) pl. XVII-XIX (1848) (37): (i) pl. XX (1849) (?38): (i) XV-XVIII (1849) (38): (i) pl. XXI-XXII (1849) (40): (ii) [I]-II-IV, pl. IV-IX (1849) (48): [1], 2-24 (1851) (55): 25-80, (iv) [1], 2-36 (1852) (60): (ii) V-VIII, pl. X-XIV (iv) 37-40 (1853) (65): (iv) 41-60 (1854) (66): (iv) 61-72 (1855) (68): 81-152 (1855) (69): 153-178, (Index univ.) [1], 2-48 (1856) (Schlusswort) [i-iv] (1856)
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816) (2): 17-32 (1819) (3): 33-48 (1819) (4): 49-64 (1819) (5): 65-80 (1819) (6): 81-96 (1819) (7): 97-112 (1819) (8): 113-128 (1819) (9): 129-144 (1819) (10): 145-160 (1819) (11): 161-176 (1819) (12): 177-192 (1820) (13): 193-208 (1820) (14): 209-224 (1821) (15): 225-240 (1821) (16): 241-256 (1821) (17): 257-272 (1823) (18): 273-288 (1823) (19): 289-304 (1823) (20): 305-320 (1825) (21): 321-336 (1825) (22): 337-352 (1825) (23-27): 353-431 ([1825])
Linnaeus, 1771
Mantissa Plantarum altera Generum editionis Vi & Specierum editionis II Mantissa Plant. 2: -,[iv], 142-510, + Regni Animalis Appendix 511-552
Oberthür, 1876
Sur la Faune des Lépidoptères de l'Algerie Étud. d'Ent. 1 : 1-74, pl. 1-4
Oberthür, 1915
Faune des Lépidoptères de la Barbarie, Algerie Étud. Lépid. Comp. 10 : 1-459, pl. 276-277,277bis,278-315
Rothschild, 1933
On a collection of Lepidoptera from Spanish Morocco Novit. Zool. 38 : 315-330
de Sagarra, 1924
Noves formes de Lepidopteres iberics Bull. Inst. catal. Hist. nat. (2) 4: 198-204
Seitz, 1906
Die Großschmetterlinge des Palaearktischen Faunengebietes. 1. Die Palaearktischen Tagfalter Gross-Schmett. Erde 1 : 1-379


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.