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Melitaea Fabricius, 1807
Melitaea Fabricius, 1807; Magazin f. Insektenk. (Illiger) 6: 284, TS: Papilio cinxia Linnaeus
Schoenis Hübner, [1819]; Verz. bek. Schmett. (2): 28, TS: Papilio delia Denis & Schiffermüller
Cinclidia Hübner, [1819]; Verz. bek. Schmett. (2): 29, TS: Papilio phoebe Denis & Schiffermüller
Melinaea Sodoffsky, 1837; Bull. Soc. imp. Nat. Moscou 10 (6): 80 (unn. repl. Melitaea Fabricius, 1807, preocc. Melinaea Hübner, 1816), TS: Papilio cinxia Linnaeus
Didymaeformis Verity, 1950; Le Farfalle diurn. d'Italia 4: 89, 90, TS: Papilio didyma Esper
Melitaea, NSG Voucher Specimen [Wahlberg]
 Melitaea (Melitaeini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165
Didymaeformia (Melitaeini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166
Cinclidia (Melitaeini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166
Melitaea (Nymphalidae) ; [BRU2, 59]
Melitaea (Melitaeina) ; [Wahlberg]; [AfrL]

The "didyma" species-group
Melitaea didyma (Esper, [1777])
NW.China. See [About maps] Spotted Fritillary Red-Band Fritillary Roter Scheckenfalter Doncella timida Przeplatka didyma
Papilio didyma Esper, [1777]; Die Schmett. Th. I, Bd. 1 (7): pl. 41 (1778), f. 3
?Papilio didyma Esper, 1781; Die Schmett. Th. I, Bd. 2 (2): pl. 61, f. 1, TL: Bavaria
Papilio cytheris Mueschen, 1781
Papilio athulia Fabricius, 1787
orientalis Herrich-Schäffer, 1856; (preocc.)
alpina Staudinger, 1861; (nom. nud.)
occidentalis Staudinger, 1861; (nom. nud.)
meridionalis Staudinger, 1861; (nom. nud.)
dalmatina Staudinger, 1861; (nom. nud.)
graeca Staudinger, 1861; (nom. nud.)
rossica Staudinger, 1861; (nom. nud.)
meridionalis Staudinger, 1870
occidentalis Staudinger, 1781
dalmatina Staudinger, 1870
Melitaea didyma, NSG Voucher Specimen [Wahlberg]
Melitaea didyma, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Melitaea didyma, Moths and Butterflies of Europe [MBE]
Melitaea didyma, Europaische Schmetterlinge [Mario Maier]
Melitaea didyma, Captain's European Butterfly Guide [Simon Coombes]
 Melitaea didyma ; [H&R, 116]
Didymaeformia didyma ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea didyma ; [BOW: pl. 3, f. 2-3]; [EBW]
Melitaea (didyma) didyma ; [BRU2: 59, pl. 38, f. 1-12, 25-26]
Melitaea didyma ; [MRS, 586]; [Otakar Kudrna]
[didyma-1fu.jpg] 500x316 (~61Kb) upperside female Axat, Pyrenees Orientales 24/07/96, Photo © Simon Coombes [didyma-1mu.jpg] 500x325 (~44Kb) upperside male Axat, Pyrenees Orientales 24/07/96, Photo © Simon Coombes [didyma-1v.jpg] 450x432 (~38Kb) underside Axat, Pyrenees Orientales 24/07/96, Photo © Simon Coombes
[didyma-2l.jpg] 597x400 (~49Kb) larva Central Europe 1990-2000, Photo © Michael Borsch [didyma-2p.jpg] 371x550 (~36Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch [didyma-2u.jpg] 597x400 (~52Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch

Larva on Plantago, Veronica, Viola, Scrophularia, Linaria, Dianthus, (etc.) [BRU2, 59]


M. d. didyma
SEU, CEU, W.Siberia, Caucasus, Transcaucasia kasanskyi Krulikovsky, 1901
marginata Ksienschopolsky, 1912; (preocc.)
niesiolowskii Bryk, 1940
uralicola Bryk, 1940
crimitaea Bryk, 1940
maledescripta Bryk, 1940
pontifex Bryk, 1940
itkolensis Alberti, 1969
Melitaea didyma didyma ; [BRU2, 59]
M. d. kirgisica Bryk, 1940
Tarbagatai, Saur, Tian-Shan ?geminella Bryk, 1940
Melitaea didyma kirgisica ; [BRU2, 59]
M. d. turkestanica Sheljuzhko, 1929
Alai Melitaea turanica Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 231 (preocc.)
pseudoala Sheljuzhko, 1928; (infrasubsp.)
Melitaea didyma turkestanica ; [BRU2, 59]
M. d. elavar Fruhstorfer, 1917
Ghissar Melitaea didyma elavar ; [BRU2, 59]
M. d. occidentalis Staudinger, 1861
Spain, N.Africa Melitaea didyma occidentalis Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8, TL: Albarracin, Spain
Melitaea didyma f. mauretanica Oberthür, 1909; Etud. Lépid. Comp. 3: 243, TL: Algeria, Spain
Melitaea didyma var. meridionalis Staudinger, 1870; Horae Soc. ent. ross. 7 (1870): 60, TL: Mt. Parnassus, Greece
Melitaea didyma occidentalis f. dalmatina Staudinger, 1870; Horae Soc. ent. ross. 7 (1870): 60, TL: Greece
Melitaea didyma nisseni Rothschild, 1913; Novit. Zool. 20 (1): 115, TL: Aïn Draham (Tunisia)
Melitaea didyma interposita Rothschild, 1913; Novit. Zool. 20 (1): 115, TL: Batna (Algeria)
Melitaea didyma mauretanica f. occasus Verity, 1919; Ent. Rec. J. Var. 31: 180, TL: [N] Africa
Melitaea didyma proteaoccidentis Verity, 1929; Ent. Rec. J. Var. 41: 117, TL: Fum Kheney [Foum Kheneg], Mekney [Meknes] (Morocco)
Melitaea didyma africa-felix Bryk, 1940; Folia zool. hydrobiol. 10: 326, pl. 1, f. 5, TL: El-Bayadh, Sebdou (Algeria)
Melitaea didyma nisseni f. gen. tunisa Eisner, 1942; Zool. Meded. 24: 118, TL: Tunis (Tunisia)
Melitaea didyma meridionalis ; [H&R, 116]
Melitaea didyma occidentalis ; [H&R, 117]; [BMAT: 53, pl. 17, f. 22-36]

Larva on Linaria spp., Scrophularia sp., Antirrhinum sp. [BMAT], (other regions) Mispate orontium, Plantago sp., Plantago amplexicaulis, Melampyrum sp., Verbascum sp., Linaria sagittata [BMAT]


interposita
[interposita-1v.jpg] 600x435 (~41Kb) underside Morocco, Tizi-n-Test, Haut Atlas, Photo © Michel Tarrier


mauretanica
[mauretanica-1uf.jpg] 600x435 (~43Kb) upperside female Morocco, Djebel Lakraa, Rif, Photo © Michel Tarrier [mauretanica-2um.jpg] 600x435 (~28Kb) upperside male Morocco, Itzer, Moyen Atlas, Photo © Jean Delacre

M. d. neera Fischer de Waldheim, 1840
Volga, S.Ural, SW.Siberia Melitaea uralca Bryk, 1940
[neera-1l.jpg] 820x480 (~92Kb) larva A high steppefied right bank of the Shipunikha Rivulet 3 km NE of the village Evsino, Iskitim district, Novosibirsk Province, West Siberia, Russia. 8th June 1996 (on a leaf of Phlomis tuberosa L.), Photo © Oleg Kosterin

Larva on Phlomis tuberosa [Oleg Kosterin]


M. d. turkestanica Sheljuzhko, 1929
?M. d. ambra Higgings, 1941
dschungarica Oberthür, 1909; (preocc.)
perplexa Higgins, 1941
manoni Hanus, 1996
ambra ; [BRU2, 59 (?)]
Melitaea interrupta Kolenati, 1846
-. See [About maps] Melitaea interrupta Kolenati, 1846; Insecta Caucasi: 86, TL: "Caucaso et Transcaucasia"
Melitaea transcaucasica Turati, 1919; Naturalista sicil. 23: 225
= ; [Wahlberg]
caucasi Verity, 1929; (preocc.)
venosata Niesiolowsky, 1937
distinctissima Bryk, 1940
musatscherii Alberti, 1969
Melitaea transcaucasica, NSG Voucher Specimen [Wahlberg]
 Didymaeformia transcaucasica ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea (didyma) interrupta ; [BRU2: 60, pl. 38, f. 13-18]
M. i. interrupta
NE.Turkey?, Caucasus, Transcaucasus Melitaea interrupta interrupta ; [BRU2, 60]
M. i. kendevana Gross & Ebert, 1975
Kopet-Dagh maculata Gross & Ebert, 1975; (preocc.)
elbursicola Gross & Ebert, 1975
safawides Gross & Ebert, 1975
?wernickei Gross & Ebert, 1975
grossi Koçak, 1980
Melitaea interrupta kendevana ; [BRU2, 60]
Melitaea latonigena Eversmann, 1847
-. See [About maps] Melitaea latonigena Eversmann, 1847; Bull. Soc. Imp. Nat. Moscou 20 (?2): 66, pl. 1, f. 1-2, TL: [Irkutsk Region, Russia]
atrata Higgins, 1935
mongoligena Bryk, 1940
Melitaea latonigena, Lepidoptera collection [SZMN]
Melitaea latonigena, NSG Voucher Specimen [Wahlberg]
 Melitaea didyma latonigena ;
Melitaea didyma latonigera[sic] ; [MRS, 586]
Melitaea (didyma) latonigena ; [BRU2: 60, pl. 37, f. 31-36]
M. l. latonigena
Tuva - Baikal Melitaea latonigena latonigena ; [BRU2, 60]
M. l. altaica Grum-Grshimailo, 1893
Altai latonigena altaica Grum-Grshimailo, 1893; , TL: SW.Altai, Lake Markakol
Melitaea latonigena altaica ; [Oleg Kosterin]; [BRU2, 60]
[altaica-1mfv.jpg] 461x700 (~58Kb) underside male female A subalpine short forb meadow on an opening in an open larch forest on the southern principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets , 2100 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 16th July 1998, Photo © Oleg Kosterin [altaica-1mv.jpg] 400x533 (~49Kb) underside male An alpine meadow, at 2200 m above sea level, on the eastern spurs of the Katunskiy Mt. Range, the valley of the Argem (Direntai) stream, Central Altai Mts., West Siberia, Russia. 14th July, 1988, Photo © Oleg Kosterin [altaica-2v.jpg] 429x668 (~28Kb) underside A larch parkland between the Chikty and Akbul Rivulets, 2000 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 15th July 1998 (on Veronica porphyriana), Photo © Oleg Kosterin
[altaica-2vx.jpg] 768x700 (~76Kb) underside A larch parkland between the Chikty and Akbul Rivulets, 2000 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 15th July 1998 (on Veronica porphyriana), Photo © Oleg Kosterin [altaica-3fu.jpg] 572x407 (~28Kb) upperside female A larch parkland between the Chikty and Akbul Rivulets, 2000 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 15th July 1998, Photo © Oleg Kosterin [altaica-3mvA.jpg] 503x432 (~21Kb) (aberrative) underside male A steppefied southern slope of small mountains on the Dzhazator River right bank in front of the Akbul Rivulet mouth, between the Yuzhno-Chuiskii mountain range and the Ukok Plateau, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 19th July 1998, Photo © Oleg Kosterin

M. l. polaris Grum-Grshimailo, 1899
Yakutia Melitaea latonigena polaris ; [BRU2, 60]
Melitaea mixta Evans, 1912
-. See [About maps] Melitaea mixta Evans, 1912; J. Bombay nat. Hist. Soc. 21 (2): 583, TL: "Chit.[ral]"
Didymaeformia mixta ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea didyma mixta ;
Melitaea (didyma) mixta ; [BRU2: 61, pl. 41, f. 1-3]
M. m. mixta
NW.India Melitaea mixta mixta ; [BRU2, 61]
M. m. nadezhdae Sheljuzhko, 1912
E.Pamirs Melitaea mixta nadezhdae ; [BRU2, 61]
M. m. problematica Sheljuzhko, 1912
W.Pamirs Melitaea mixta problematica ; [BRU2, 61]
Melitaea persea Kollar, [1850]
Iran, Asia Minor - Afghanistan, W.Tien-Shan. See [About maps] Melitaea persea Kollar, [1850]; Denkschr. Akad. Wiss. Wien. 1: 11, TL: "Farsistan" [Shiraz, Iran]
kaschtschenkoi Christoph, 1889
= ; [BRU2, 61]
Melitaea didyma magnacasta Verity, 1929
= ; [BRU2, 61]
Melitaea didyma araratica Verity, 1929
= ; [BRU2, 61]
Melitaea didyma tauricus Belter, 1934
= ; [BRU2, 61]
microtauricus Belter, 1934
= ; [BRU2, 61]
hafiz Higgins, 1941
= ; [BRU2, 61]
afghana Heydemann, 1954
= ; [BRU2, 61]
darius Gross & Ebert, 1975
= ; [BRU2, 61]
pfeifferi Gross & Ebert, 1975
= ; [BRU2, 61]
?wernickei Gross & Ebert, 1975
= ; [BRU2, 61]
Melitaea persea, NSG Voucher Specimen [Wahlberg]
 Didymaeformia afghana ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Didymaeformia persea ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea persea ; Weiss & Major, 2000, Atalanta 31 (1/2): 123 (note), pl. VII, f. 9-16; [AfrL]; Carbonell, 2007, Bull. Soc. ent. Fr. 112 (3): 334 (note)
Melitaea (didyma) persea ; [BRU2: 61, pl. 44, f. 1-9]
M. p. caucasica Staudinger, 1870
caucasica Staudinger, 1861; (nom. nud.)
M. p. dogsoni Grose-Smith, 1887
M. p. paphlagonia Fruhstorfer, 1917
M. p. montium Belter, 1934
Syria, Lebanon Melitaea montium Belter, 1934; Arb. Morph. Tax. Ent. Berl. 1: 107
= Melitaea persea Kollar, [1850]; [BRU2, 61]
M. p. sargon Hemming, 1932
Melitaea persea sargon ; [AfrL]
Melitaea jitka Weiss & Major, 2000
SW.Iran. See [About maps] Melitaea jitka Weiss & Major, 2000; Atalanta 31 (1/2): 123, pl. V, f. 1-4, pl. VI, pl. VII, f. 1-8, TL: SW.Iran,Buyer Ahmed-Kuhgilueye, Yasuj, Dinar Mts., 2600-2700m
Melitaea casta (Kollar, [1848])
Iran. See [About maps] Melitatea casta Kollar, [1848]; Denkschr. Akad. Wiss. Wien. 1: 52
Melitaea casta, NSG Voucher Specimen [Wahlberg]
 Didymaeformia casta ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea casta ; Weiss & Major, 2000, Atalanta 31 (1/2): 126 (note), pl. V, f. 5-8
Melitaea saxatilis Christoff, 1876[1873?]
Iran. See [About maps] Melitaea saxatilis Christoff, 1876[1873?]; Hor. Soc. Ent. Ross. 10: 28
Melitaea saxatilis, NSG Voucher Specimen [Wahlberg]
 Didymaeformia saxatilis ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea saxatilis ; [BOW: pl. 196, f. 7]
Melitaea deserticola Oberthür, 1909
N.Africa, Tripolitania, Egypt, Lebanon, Transjordan. See [About maps] Desert Fritillary
Melitaea ab. deserticola Oberthür, 1876; Etud. Ent. 1: 25, pl. 3, f. 1, TL: Biskra (Algeria)
Melitaea deserticola, NSG Voucher Specimen [Wahlberg]
Melitaea deserticola, Moths and Butterflies of Europe [MBE]
 Melitaea deserticola ; [H&R, 117]
Didymaeformia deserticola ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea deserticola ; [AfrL]
[deserticola-1u.jpg] 600x435 (~35Kb) upperside Morocco, Plateau de l'Arid, Haut Atlas, Photo © Jean Delacre [deserticola-2uf.jpg] 600x435 (~68Kb) upperside female Morocco, Plateau de l'Arid, Haut Atlas, Photo © Jean Delacre [deserticola-3v.jpg] 600x435 (~59Kb) underside Morocco, Plateau de l'Arid, Haut Atlas, Photo © Jean Delacre
[deserticola-4u.jpg] 600x435 (~53Kb) upperside Morocco, Aït-Abdallah, Anti-Atlas , Photo © Michel Tarrier

M. d. deserticola
Tunisia, Algeria, Morocco Melitaea didyma harterti Rothschild, 1913; Novit. Zool. 20 (1): 115, TL: El Hadadra, El Golea, Ghardaia (Algeria)
Melitaea didyma harterti ab. pseudoacraeina Rothschild, 1917; Novit. zool. 24: 102, TL: El Hadadra (Algeria)
Melitaea didyma harterti ab. nigrofasciata Rothschild, 1917; Novit. zool. 24: 102, TL: El Hadadra (Algeria)
Melitaea deserticola deserticola ; [BMAT: 54, pl. 17, f. 37-38, pl. 18, f. 1-7]

Larva on Linaria aegyptiaca, Plantago media, Anarrhinum fruticosum, (in Lebanon) Anarrhinum spp. [BMAT]

M. d. macromaculata Belter, 1934
Melitaea deserticola macromaculata ; [AfrL]
M. d. scotti Higgins, 1941
Melitaea abyssinica scotti Higgins, 1941
Melitaea deserticola scotti ; [AfrL]
Melitaea abyssinica Oberthür, 1909
Ethiopia. See [About maps] Didymaeformia abyssinica ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea abyssinica ; [BAFR, 213]; [AfrL]
Melitaea didymina Staudinger, 1895
Mongolia. See [About maps] Melitaea didymina Staudinger, 1895; Dt. Ent. Z. Iris 8: 347
Didymaeformia didymina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea didymoides Eversmann, 1847
-. See [About maps] Melitaea didymoides Eversmann, 1847; Bull. Soc. Imp. Nat. Moscou 20 (?2): 67, pl. 1, f. 3-4, TL: [Kyakhta, Buryatia]
sibirica Staudinger, 1861; (nom. nud.)
sibirica [Heyne], [1893]; (preocc.)
mandschurica Seitz, [1909]; (preocc.)
seitzi Matsumura, 1929
mandschukoana Bryk, 1940
Melitaea didymoides, NSG Voucher Specimen [Wahlberg]
 Didymaeformia didymoides ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea didymoides ; [MRS, 586]; [BRU2: 62, pl. 39, f. 19-24]
[didymoides-1mv.jpg] 592x710 (~76Kb) underside male A steppefied meadow at Lake Butyvken, surrounded by a magnificent pine forest within a steppe zone called Tsasucheiskii Bor, 10 km SW of the village Nizhnii Tsasuchei, Onon District, SE Chita Province, Transbaikalia, Siberia, Russia. 4th July 1996, Photo © Oleg Kosterin

M. d. didymoides
S.Tuva?, Transbaikalia, Amur Melitaea didymoides didymoides ; [BRU2, 62]
M. d. yagakuana Matsumura, 1927
S.Ussuri Melitaea didymoides yagakuana ; [BRU2, 62]
M. d. pekinensis Seitz, [1909]
N.China Melitaea didymoides pekinensis ; [MRS, 586]; [BRU2, 63]
M. d. latonia Grum-Grshimailo, 1891
C.China Melitaea didymoides latonia ; [MRS, 586]; [BRU2, 62]
M. d. eupatides Fruhstorfer, 1917
Gansu cansicola Bryk, 1940
Melitaea didymoides eupatides ; [BRU2, 63]
M. d. hummeli Bryk, 1940
S.Mongolia Melitaea didymoides hummeli ; [BRU2, 63]
Melitaea ala Staudinger, 1881
Tian-Shan, Dzhungarsky Ala-Tau Mts., S.Altai. See [About maps] Melitaea didyma var. ala Staudinger, 1881; Stett. ent. Ztg 42: 288, TL: Lepsa, Ala-Tau
Melitaea ala, NSG Voucher Specimen [Wahlberg]
 Didymaeformia ala ; Higgings, 1981, : 166 (name)
Melitaea ala ; [BRU2: 63, pl. 39, f. 10-15]

Larva on Ligularia [BRU2, 63]

M. a. ala
S.Altai, Saur Mts., Dzhungarsky Ala-Tau, Burchansarytau Mts., Tyshkantau Mts., Kaiandy-Tau, Horgos R., Alty-Emel Mts. Melitaea ala ab. immodulata Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 366, TL: Dzharkent
Melitaea ala allah Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 346, pl. 3, f. 23, TL: "Siberia"
Melitaea ala ; Sushkin, 1913, Z. wiss. Insektenbiologie 9 (6/7): 171, f. 5-6; Wagner, 1913, Entomol. Mitteilungen 2 (5): 91-93
Melitaea ala ala ; Sheljuzhko, 1929, Mitt. Münch. Ent. Ges. 19: 368; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 346
Melitaea ala ; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 242-244, pl. 2, f. 4, 10; Lukhtanov & Lukhtanov, 1994, Herbipoliana, 3: 184-185, pl. 36, f. 5-6
Melitaea ala ala ; Kolesnichenko, 1999, Atalanta 30 (1/4): 91, pl. 6, f. 1-4; [BRU2, 63]
M. a. bicolor Seitz, 1907
Kungey Ala-Tau, Kirgizsky Mts., Suusamyr Valley, Suusamyr Mts., Talassky Ala-Tau, Terskey Ala-Tau, Baidullu Mts., Naryn-Too Mts. Melitaea ala ab. bicolor Seitz, 1907; Gross-Schmett. Erde 1: 219, pl. 66, f. 7-8, TL: Karagatai Mts.
Melitaea ala ab. latemarginata Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 366-367, pl. 28, f. 7-8, TL: Naryn
Melitaea ala bicolor ; Sushkin, 1913, Z. wiss. Insektenbiologie 9 (6/7): 172; Sheljuzhko, 1929, Mitt. Münch. Ent. Ges. 19: 368; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 347-348; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 244, pl. 6, f. 6, 12; Kolesnichenko, 1999, Atalanta 30 (1/4): 93, pl. 6, f. 5-7; [BRU2, 63]
M. a. zaisana Lukhtanov, 1999
S.Altai Melitaea ala zaisana Lukhtanov, 1999; Atalanta 30 (1/4): 143, TL: Kazakstan, Ust-Kamenogorsk, S.Altai, between Kurtschum and Kalguty, 600m
Misidentified/misapplied:
Melitaea ala; Lukhtanov & Lukhtanov, 1994, Herbipoliana, 3: pl. 35, f. 5-6
M. a. irtyschica Lukhtanov, 1999
W.Altai Melitaea ala irtyschica Lukhtanov, 1999; Atalanta 30 (1/4): 144, TL: Kazakstan, Ust-Kamenogorsk, Ulbinski, Oktjabrsk, 420m
?M. a. sheljuzhkoi Bryk, 1940
E.Tian-Shan Melitaea ala sheljuzhkoi ; [BRU2, 63 (?)]
?M. a. determinata Bryk, 1940
E.Tian-Shan Melitaea ala determinata ; [BRU2, 63 (?)]
?M. a. strandi Bryk, 1940
E.Tian-Shan Melitaea ala strandi ; [BRU2, 63 (?)]
Melitaea kotshubeji Sheljuzhko, 1929
Peter the Great Mts., N.Hazreti-Sho Mts., W.Zaalaisky Mts., S slopes (Alaisky Mts.), SW slopes (Fergansky Mts.), SE slopes (Cathkalsky Mts.) ?kotshubei; (spell.?)
Melitaea kotshubeji ; [BRU2: 64, pl. 38, f. 31-32, pl. 39, f. 1-3]

Larva on Ligularia thomsonii [BRU2, 65]

M. k. kotshubeji
Peter the Great Mts., Hazreti-Sho, Zaalaisky Mts. Melitaea didyma var. ala Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4: 430, TL: Touptshek
Melitaea ala kotshubeji Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 364, pl. 27, f. 1-4, TL: Touptshek
Melitaea ala kotshubeji ; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 348
Melitaea chitralensis mod. kosthubeji ; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 248, pl. 6, f. 4, 10
Melitaea ala kotshubeji ; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 48
Melitaea kotshubeji kotshubeji ; Kolesnichenko, 1999, Atalanta 30 (1/4): 96, pl. 6, f. 8-11
Misidentified/misapplied:
Melitaea ala bicolor Seitz; Shchetkin, 1981, Ent. Obozr. Tajikistana: 163-164

Larva on Ligularia thompsonii Shchetkin, 1981, Ent. Obozr. Tajikistana

M. k. bundeli Kolesnichenko, 1999
W.Zaalaisky Mts., S.Alaisky Mts. Melitaea kotshubeji bundeli Kolesnichenko, 1999; Atalanta 30 (1/4): 98, pl. 6, f. 12-13, TL: Alaisky Mts., near Daraut-Kurgan, 3000
Melitaea kotshubeji bundeli ; [BRU2, 65]
M. k. kugarti Kolesnichenko, 1999
northern slopes (E.Alaisky Mts.), Fergansky, Chatkalsky Mts. Melitaea kotshubeji kugarti Kolesnichenko, 1999; Atalanta 30 (1/4): 100, pl. 6, f. 14-17, TL: Fergansky Mts., Kugart river, 2000
Melitaea ninae Sheljuzhko, 1935
Chatkalsky Mts., Pskempsky Mts., Kuraminsky Mts., Talassky Ala-Tau Mts., Vissokoe, Chandalakshsky Mts. Melitaea ala ninae Sheljuzhko, 1935; Mitt. Münch. ent. Ges. 25: 28, TL: Tian-Shan, Tshimgan
Melitaea ala ninae f. magnifica Sheljuzhko, 1935; Mitt. Münch. ent. Ges. 25: pl. 3, f. 5
Melitaea ala ninae f. oblongomaculata Sheljuzhko, 1935; Mitt. Münch. ent. Ges. 25: pl. 3, f. 6
Melitaea ala ninae f. elongatoconfluens Sheljuzhko, 1935; Mitt. Münch. ent. Ges. 25:
Melitaea rosea Higgins, 1938; Entomologist 71: 34, TL: Alie-Ata
Melitaea ala ella-claudia Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 351, pl. 5, f. 49-50, pl. 6, f. 54, TL: Aulie-Ata, Talas Alatau
Melitaea ninae, NSG Voucher Specimen [Wahlberg]
 elongotoconfluens ; [BRU2, 64 (missp.?)]
Melitaea ninae ; Lukhtanov & Lukhtanov, 1994, Herbipoliana, 3: 85, pl. 36, f. 7-8; Kolesnichenko, 1999, Atalanta 30 (1/4): 101, pl. 7, f. 18-21; [BRU2: 64, pl. 39, f. 7-9]
Misidentified/misapplied:
Melitaea pseudoala Sheljuzhko; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 245, pl. 4, f. 12, pl. 16, f. 4; Higgins, 1955, Trans. R. ent. Soc. Lond. 106 (1): 116; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166; Korshunov, 1972, Ent. Obozr. 51 (2): 355; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 48
Melitaea chitralensis Moore, 1901
Kashmir, Chitral. See [About maps] Melitaea chitralensis Moore, 1901; Lepidoptera Indica, 5: 9, pl. 380, f. 1, TL: Chitral
Melitaea didyma chitralipluvia Verity, 1929; Ent. Rec. J. Var. 41: 43, TL: Chitral
Melitaea chitralensis, NSG Voucher Specimen [Wahlberg]
 Melitaea chitralensis ; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 247, pl. 6, f. 3, 9; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name); Kolesnichenko, 1999, Atalanta 30 (1/4): 103, pl. 7, f. 22-25
Melitaea enarea Fruhstorfer, 1917
Melitaea didyma enarea Fruhstorfer, 1917; Archiv Naturgesch. (A) 82 (2): 11, TL: Garm, Peter the Great Mts.
Melitaea enarea, NSG Voucher Specimen [Wahlberg]
 Melitaea enarea ; [BRU2: 64, pl. 38, f. 19-24, 27-30]
M. e. enarea
Garm, Kamara-su river, Peter the Great Mts., Hazreti-Sho, Darvazky Mts., Shugnansky Mts. Melitaea didyma shugnana Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 355, pl. 27, f. 7-10, TL: Chorag

Melitaea didyma enarea ; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 333, pl. 1, f. 7, pl. 5, f. 52
Melitaea chitralensis enarea ; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 248, pl. 6, f. 2; Shchetkin, 1981, Ent. Obozr. Tajikistana: 164-165
 Melitaea didyma enarea ; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 47
Melitaea chitralensis shugnana ; Tshikolovets, 1997, The Butterflies of Pamir: 118, pl. 28, f. 7-8, pl. 29, f. 7-8, pl. 43, f. 4
Melitaea enarea enarea ; Kolesnichenko, 1999, Atalanta 30 (1/4): 106, pl. 7, f. 26-29
Melitaea enarea shugnana ; [BRU2, 64]
Melitaea enarea enarea ; [BRU2, 64]
M. e. ishkashima Sheljuzhko, 1929
Ishkashimsky Mts. Melitaea didyma ishkashima Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 359, pl. 27, f. 13-14, TL: Kishlak Njut
= Melitaea enarea shugnana Sheljuzhko, 1929; [BRU2, 64]
Melitaea didyma ishkashima ; Tshikolovets, 1997, The Butterflies of Pamir: pl. 28, f. 13, pl. 29, f. 13, pl. 38, f. 8, pl. 39, f. 8
Melitaea enarea ishkashima ; Kolesnichenko, 1999, Atalanta 30 (1/4): 108, pl. 7, f. 30
M. e. permuta Higgins, 1941
Nuratau Mts., Kugitang Mts., Gissarsky Mts., Zeravshansky Mts., Turkenstansky Mts., Karateginsky Mts. Melitaea enarea mod. permuta Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 250, pl. 6, f. 7-8, TL: Samarkand, Hazreth-Sultan
Melitaea permuta, NSG Voucher Specimen [Wahlberg]
 Melitaea ala permuta ; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 48
Melitaea enarea permuta ; Kolesnichenko, 1999, Atalanta 30 (1/4): 109, pl. 7, f. 31
Melitaea permuta ; [BRU2: 64, pl. 39, f. 4-6]
M. e. gromenkoi Kolesnichenko, 1999
Fergansky Mts. Melitaea enarea gromenkoi Kolesnichenko, 1999; Atalanta 30 (1/4): 110, pl. 7, f. 32-34, TL: Kirghizia, 15km NW from Dzhalal-Abad, Kara-Kianyr Mts., 1200m
Melitaea enarea gromenkoi ; [BRU2, 64]
Melitaea pseudoala Sheljuzhko, 1928
Turkestan. See [About maps] Melitaea pseudoala Sheljuzhko, 1928; : 355
= Melitaea didyma turkestanica Sheljuzhko, 1928; [Wahlberg]
Didymaeformia pseudoala ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea sutschana Staudinger, 1892
-. See [About maps] Melitaea sutschana Staudinger, 1892; in Romanoff, Mém. Lépid., 6: 183, TL: Sutschan-Gebiete
sutschanala Bryk, 1940
transbaicalica Bryk, 1940
Melitaea sutschana, NSG Voucher Specimen [Wahlberg]
 Didymaeformia sutschana ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea sutschana ; [BRU2: 63, pl. 39, f. 16-18]
[sutschana-1fv.jpg] 410x400 (~37Kb) underside female A steppefied meadow on a southern slope of the Polovinnaya Pad' valley few kilometres of the Argun' River left bank 13 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 28th July 1997 (aberrantly large basal black spots), Photo © Oleg Kosterin [sutschana-2fu.jpg] 410x547 (~53Kb) upperside female A steppefied meadow on the Argun' River left bank terrace 4 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 26th July 1997, Photo © Oleg Kosterin [sutschana-3mv.jpg] 533x400 (~35Kb) underside male A steppefied southern left slope of the Budyumkan River valley 5 km of its confluence with the Argun' River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 26th July 1997 (on Leontopodium conglobatum (Turcz.) Hand.-Mazz.), Photo © Oleg Kosterin
[sutschana-4mu.jpg] 565x490 (~34Kb) upperside male A steppefied southern left slope of the Budyumkan River valley 5 km of its confluence with the Argun River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 26th July 1997 (on Leontopodium conglobatum (Turcz.) Hand.-Mazz), Photo © Oleg Kosterin

M. s. sutschana
Transbaikalia - Amur, Ussuri, Korea, NE.China Melitaea sutschana sutschana ; [BRU2, 63]
M. s. graeseri Gorbunov, 1995
Sakhalin Melitaea sutschana graeseri ; [BRU2, 63]
Melitaea acraeina (Staudinger, 1886)
Fergana Valley Melitaena acraeina Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 233, TL: Kokand and Fergana, Fergana Valley, Uzbekistan
Melitaea acraeina, NSG Voucher Specimen [Wahlberg]
 Didymaeformia acraeina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea acraeina ; [BOW: pl. 195, f. 4]; [BRU2: 65, pl. 44, f. 13-15]

Larva on Dodartia orientalis [BRU2, 65]

Melitaea lunulata Staudinger, 1901
-. See [About maps] Melitaea lunulata Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 1: 30, TL: Issyk-Kul
Melitaea lunulata, NSG Voucher Specimen [Wahlberg]
 Didymaeformia lunulata ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea lunulata ; [BRU2: 65, pl. 43, f. 16-18]
M. l. lunulata
Tian-Shan Melitaea lunulata lunulata ; [BRU2, 66]
M. l. merke Lukhtanov, 1999
W.Tian-Shan Melitaea lunulata merke Lukhtanov, 1999; Atalanta 30 (1/4): 147, TL: Kazakstan, Merke Fl., 2900
Melitaea lunulata merke ; [BRU2, 66]
Misidentified/misapplied:
Melitaea fergana lunulata; Lukhtanov & Lukhtanov, 1994, Herbipoliana, 3: pl. 33, f. 5
Melitaea fergana Staudinger, 1882
Turkestan. See [About maps] Melitaea fergana Staudinger, 1882; Berl. Ent. Z. 26: 168, TL: Alai-Gebirge [Fergana, Turkestan]
Melitaea fergana, NSG Voucher Specimen [Wahlberg]
 Didymaeformia fergana ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea fergana ; [BRU2: 66, pl. 42, f. 31-42, pl. 43, f. 1-12]
M. f. fergana
Alai, Transalai Melitaea fergana fergana ; [BRU2, 66]
M. f. paradoxa Churkin & Tuzov, 2000
N.Tian-Shan Melitaea fergana paradoxa Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
Melitaea fergana paradoxa ; [BRU2, 66]
M. f. ketmeana Lukhtanov, 1999
S.Kazakstan, NE.Kirghizia Melitaea fergana ketmeana Lukhtanov, 1999; Atalanta 30 (1/4): 145, TL: Kazakstan, Alma-Ata, Ketmen, Kolshat
Melitaea fergana ketmeana ; [BRU2, 66]
M. f. terskeana Lukhtanov, 1999
Kirghizia Melitaea fergana terskeana Lukhtanov, 1999; Atalanta 30 (1/4): 146, TL: Kirghizia, Terskey-Alatoo, Barskoun-Pass, 3400m
Melitaea fergana terskeana ; [BRU2, 66 (?)]
M. f. oxuana Lukhtanov, 1999
E.Pamir Melitaea fergana oxuana Lukhtanov, 1999; Atalanta 30 (1/4): 147, TL: Tadjikistan, E.Pamir, Sarykol, Dunkeidyk, 3750
M. f. grumi Churkin & Tuzov, 2000
E.Pamir Melitaea fergana grumi Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
Melitaea fergana grumi ; [BRU2, 66]
M. f. vladislavi Churkin & Tuzov, 2000
Inner Tian-Shan (At-Bashi Mts.) Melitaea fergana vladislavi Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
Melitaea fergana vladislavi ; [BRU2, 66]
M. f. khantengri Churkin & Tuzov, 2000
Inner Tian-Shan (Kaindy Mts.) Melitaea fergana khantengri Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
Melitaea fergana khantengri ; [BRU2, 66]
M. f. maracandica Staudinger, 1882
Ghissarsky Mts. Melitaea maracandica Staudinger, 1882; Berl. Ent. Z. 26: 166
Melitaea maracandica, NSG Voucher Specimen [Wahlberg]
 Didymaeformia macarandica[sic] ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name) [spell.?]
Melitaea fergana maracandica ; [BRU2, 66]
M. f. splendida Churkin & Tuzov, 2000
Turkestansky Mts. (Pass Kumbel) Melitaea fergana splendida Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
M. f. darvasika Higgins, 1941
Darvaz Melitaea fergana darvasika ; [BRU2, 66]
M. f. jacobsoni Higgins, 1941
W.Pamirs Melitaea fergana jacobsoni ; [BRU2, 66]
Melitaea infernalis Grum-Grshimailo, 1891
Tian-Shan, Dzhungarsky Alatau. See [About maps] Melitaea infernalis Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 455
Melitaea infernalis, NSG Voucher Specimen [Wahlberg]
 Didymaeformia infernalis ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea infernalis ; [BRU2: 67, pl. 43, f. 13-15]
Melitaea ambrisia Higgins, 1935
Alai, Ghissar. See [About maps] Melitaea ambrisia Higgins, 1935; Entomologist 68: 112, TL: Alai Mts., Kirgizia
Melitaea ambrisia, NSG Voucher Specimen [Wahlberg]
 Didymaeformia ambrisia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea ambrisia ; Churkin, 2000, Atalanta 31 (1/2): 114, pl. IV, f. 1-3; [BRU2: 67, pl. 43, f. 25-30]
M. a. ambrisia
Alai Melitaea ambrisia ambrisia ; [BRU2, 67]
M. a. avicaena Churkin, 2000
Turkestansky Mts. (Kumbel Pass) Melitaea ambrisia avicaena Churkin, 2000; Atalanta 31 (1/2): 115, pl. IV, f. 4-6, TL: Uzbekistan, Turkestansky Range, Kumbel Pass, Zaaminsky rs., 3000m
Melitaea ambrisia avicaena ; [BRU2, 67]
Melitaea alraschid Higgins, 1941
-. See [About maps] Melitaea ambrisia alraschid Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 259-260, TL: Persia [= error] [?W Gissar, near Sarmarkand]
Melitaea alraschid ; Churkin, 2000, Atalanta 31 (1/2): 119, pl. IV, f. 7-9; [BRU2: 67, pl. 43, f. 19-24]
M. a. alraschid
Ghissar Melitaea alraschid alraschid ; [BRU2, 67]
M. a. garun Churkin, 2000
Matcha Melitaea alraschid garun ; [BRU2, 67]
Melitaea shandura Evans, 1924
Afghanistan, Pamirs, Pakistan, NW.India. See [About maps] Melitaea shandura Evans, 1924; J. Bombay nat. Hist. Soc. 30: 90, TL: Shandur, Chitral [NW.India]
shandura Evans, 1912; (nom. nud.)
Melitaea shandura, NSG Voucher Specimen [Wahlberg]
 Didymaeformia shandura ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea shandura ; [BRU2: 68, pl. 43, f. 31-33]
M. s. pavlitzkajana Sheljuzhko, 1943
W.Pamirs Melitaea shandura apsara Wyatt, 1961; J. Lep. Soc. 15 (1): 6, TL: above Kars Pass, Shiva Mts, N. Badakhshan, 12500-14000ft, Afghanistan
Melitaea shandura pavlitzkajana ; [BRU2, 68]
Melitaea athene Staudinger, 1881
Saur, Tarbagatai Melitaea athene Staudinger, 1881; Stett. ent. Ztg 42: 266, TL: Saisan [Saur Mts. near Zaisan, E.Kazakhstan]
Melitaea athene, NSG Voucher Specimen [Wahlberg]
 Didymaeformia athene ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea athene ; [BRU2: 65, pl. 43, f. 34-36]
Melitaea trivia ([Schiffermüller], 1775)
-. See [About maps] Lesser Spotted Fritillary Desert Fritillary Braunlicher Scheckenfalter
Papilio trivia [Schiffermüller], 1775; Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend: 179, TL: Vienna
Papilio iphigenia Esper, 1782; (preocc.)
Papilio antigonus Herbst, 1800
cleo (Latreille, 1803)
Melitaea trivia interrupta Skala, 1907
Melitaea didyma aabaca Fruhstorfer, 1917
Melitaea trivia lathon Fruhstorfer, 1917
Melitaea catapeloides Stauder, 1918
Melitaea trivia tigris Higgins, 1941
Melitaea trivia, NSG Voucher Specimen [Wahlberg]
Melitaea trivia, Moths and Butterflies of Europe [MBE]
Melitaea trivia, Europaische Schmetterlinge [Mario Maier]
 Didymaeformia trivia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea trivia ; [H&R, 118]; [BOW: pl. 3, f. 5]; [EBW]; [BRU2: 62, pl. 40, f. 1-36]; [Otakar Kudrna]

Larva on Verbascum thapsus [H&R]

M. t. fascelis (Esper, [1783])
S.Europe
Papilio fascelis Esper, [1783]
minor Ksienschopolsky, 1912
 striata Ksienschopolsky, 1912
paula Obraztsov, 1936
Melitaeta trivia fascelis ; [BRU2, 62]

Larva on Scrophularia, Eremostachys [BRU2, 62]

M. t. uvarovi Gorbunov, 1995
Kazakhstan, SW.Siberia, Tarbagatai, Saur, Dzhungarsky Alatau Melitaea trivia uvarovi ; [BRU2, 62]
M. t. singularia Korshunov, 1995
Tuva Melitaea trivia singularia, Lepidoptera collection [SZMN]
 Melitaea trivia singularia ; [BRU2, 62]
M. t. caucasi Verity, 1922
Caucasus, Transcausia Melitaea trivia caucasi ; [BRU2, 62]
M. t. nana Staudinger, 1871
Kopet-Dagh
pseudodidyma Rebel, 1905
syriaca Rebel, 1905
 aabaca Fruhstorfer, 1917
wyatti Gross & Ebert, 1975
Melitaea trivia nana ; [BRU2, 62]
M. t. catapelia Staudinger, 1886
Ghissar, Alai, Tian-Shan Melitaea trivia bactriana Shchshetkin, 1984
= ; [BRU2, 62]
Melitaea trivia catapelia ; [BRU2, 62]
M. t. chorosana Shchetkin, 1984
S.Ghissar Melitaea trivia chorosana ; [BRU2, 62]
M. t. petri Shchetkin, 1984
Darvaz Melitaea trivia petri ; [BRU2, 62]
M. t. nativa Tuzov, 2000
W.Pamirs Melitaea trivia nativa Tuzov, 2000; [BRU2, 62 (repl.)]
Melitaea catapelia shugnana Shchetkin, 1984; (preocc. Melitaea didyma shugnana Sheljuzhko, 1929)
M. t. ignasiti Sagarra, 1926²
Portugal trivia ignasiti Sagarra, 1926²; , TL: Portugal
Melitaea trivia ignasiti ; [H&R, 118]
Melitaea robertsi Butler, 1880
Baluchistan, Chitral. See [About maps] Baluchi Fritillary
Melitaea yuenty Oberthür, 1888
W.China. See [About maps] Melitaea yuenty Oberthür, 1888; Etudes d'Ent. 11: 17, TL: Tatsienlu
Didymaeformia yuenty ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea yuenty ; [BOW: pl. 196, f. 9]; [MRS, 587]
M. y. yuenty
M. y. batangensis Belter, 1944
Melitaea yuenty batangensis Belter, 1944; , TL: Batang
Melitaea yuenty batangensis ; Huang, 2001
Melitaea lutko Evans, 1932
W.Himalayas. See [About maps] Melitaea lutko Evans, 1932; Indian Butterflies (edn. 2): 185
Melitaea lutko, NSG Voucher Specimen [Wahlberg]
 Melitaea lutko ; [BOW: pl. 196, f. 4]
?Melitaea lukto[sic] ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name) [spell.?]
Melitaea mimetica Higgins, 1940
Pakistan, Afghanistan, Turkmenia. See [About maps] Melitaea mimetica Higgins, 1940; Entomologist 73: 52, TL: Baluchistan
= ; [Wahlberg]
Melitaea mimetica ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name); [BRU2: 61, pl. 44, f. 10-12]
Melitaea romanovi Grum-Grshimailo, 1891
N.China, Mongolia, Transbaikalia. See [About maps] Melitaea romanovi Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 454, TL: [Gansu, China]
Melitaea romanovi, NSG Voucher Specimen [Wahlberg]
 Melitaea romanovi ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name); [MRS, 587]; [BRU2: 65, pl. 44, f. 16-18]
M. r. romanovi
N.China Melitaea romanovi romanovi ; [BRU2, 65]
M. r. puella Higgins, 1941
Transbaikalia Melitaea romanovi puella, Lepidoptera collection [SZMN]
 Melitaea romanovi puella ; [Oleg Kosterin]; [BRU2, 65]
[puella-1fv.jpg] 800x600 (~103Kb) underside female The southern slope, covered with a short-grass steppe, of the hill Chikhalan on the northern bank of a brackish Lake Zun-Torei, Onon district, Chita Province, S Transbaikalia (Dahuria), Siberia, Russia. 12th July 1996, Photo © Oleg Kosterin [puella-2f.jpg] 600x467 (~24Kb) female A short-grass steppe, on the northern bank of a brackish Lake Zun-Torei at the foot of the hill Chikhalan, Onon district, Chita Province, S Transbaikalia (Dahuria), Siberia, Russia. 13th July 1996 (captured by a spider), Photo © Oleg Kosterin

Melitaea arduinna (Esper, 1783)
SEEU, S.Russia, Asia Minor, Iran, C.Asia. See [About maps] Freyer's Fritillary
Papilio arduinna Esper, 1783; Die Schmett. Th. I, Bd. 2 (8): 169, pl. 87, f. 4, TL: [Kamyshin, Volgograd Region, Russia]
Melitaea arduinna, NSG Voucher Specimen [Wahlberg]
 Melitaea arduinna ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [BOW: pl. 2, f. 28 (text only)]; [H&R, 114]; [BRU2: 72, pl. 45, f. 1-9]; [Otakar Kudrna]

Larva on Centaurea behen, (Transcaucasia) [BRU2, 72]

M. a. arduinna
SEEU, SW.Siberia, Altai, Alai, N.Tian-Shan, Dzhungarsky Alatau, Tarbagatai, Saur Melitaea arduinna arduinna ; [BRU2, 72]
M. a. uralensis Eversmann, 1844
arduinna uralensis Eversmann, 1844; , TL: S.Urals
Melitaea arduinna uralensis ; [BRU2, 72]
M. a. rhodopensis Freyer, [1836]
Melitaea rhodopensis Freyer, [1836]; Neuere Beitr. Schmett. 3 : 3, pl. 193, f. 1, TL: ?Macedonia "europäischen Türkei"
rhodopensis ; Abadjiev, 2002, Neue Ent. Nachr. 23: 45
Melitaea arduinna rhodopensis ; [H&R, 114]
M. a. kocaki Wagener & Gross, 1976
Transcaucasis Melitaea arduinna kocaki ; [BRU2, 72]
M. a. evanescens Staudinger, 1886
Kopet-Dagh, Ghissar-Darvaz, W.Pamirs Melitaea arduinna evanescens ; [BRU2, 72]
Melitaea avinovi Sheljuzhko, 1914
Afghanistan, W.Pamirs. See [About maps] Melitaea avinovi Sheljuzhko, 1914; Dt. Ent. Z. Iris 28: 20, TL: [Nyut, Ishkashimsky Mts., W.Pamirs, Tajikistan]
Melitaea avinovi, NSG Voucher Specimen [Wahlberg]
 Melitaea avinovi ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [BRU2: 74, pl. 44, f. 34-36]
Melitaea agar Oberthür, 1888
Tibet. See [About maps] Melitaea agar Oberthür, 1888; Etudes d'Ent. 11: 18
Melitaea agar Oberthür, 1886
Melitaea didyma wardi Watkins, 1927
Melitaea didyma baileyi Watkins, 1927
Melitaea agar ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [BOW: pl. 196, f. 2]; [MRS, 587]
Melitaea agar baileyi ; [MRS, 587]
Melitaea sarvistana Wiltshire, 1941
Iran. See [About maps] Melitaea sarvistana Wiltshire, 1941; J. Bomb. Nat. Hist. Soc. 42 (3): 473
Melitaea sarvistana, NSG Voucher Specimen [Wahlberg]
 Cinclidia sarvistana ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea sarvistana ; Carbonell, 2007, Bull. Soc. ent. Fr. 112 (3): 334 (note)
Melitaea meherparvari Carbonell, 2007
Iran. See [About maps] Melitaea meherparvari Carbonell, 2007; Bull. Soc. ent. Fr. 112 (3): 334, TL: O. Iran, "ca. Mamulan, 900-950m, Lorestan

The "cinxia" species-group
fiMelitaea cinxia (Linnaeus, 1758)
EU, Morocco, Russia, W.Asia, Amurland. See [About maps] Täpläverkkoperhonen Glanville Fritillary Hökblomsternätfjäril Pricknätfjäril Wegerich-Scheckenfalter Le Damier Doncella punteada Przeplatka cinksia
Papilio cinxia Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 480, TL: Sweden
Papilio pilodellae Rottemburg, 1775
Papilio pilosella Rottemburg, 1775
Papilio delia Denis & Schiffermüller, 1775; Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend: 179
Papilio abacus Rezius, 1783; (nom. nud.)
Papilio cinxia var. fulla Quensel, 1791
horvathi Abafi-Aigner, 1906
kasyi Gross & Ebert, 1975
Melitaea cinxia, NSG Voucher Specimen [Wahlberg]
Melitaea cinxia, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Melitaea cinxia, Svenska fjärilar [SVF]
Melitaea cinxia, Norges Sommerfugler [NORLEP]
Melitaea cinxia, Moths and Butterflies of Europe [MBE]
Melitaea cinxia, Europaische Schmetterlinge [Mario Maier]
 Melitaea cinxia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [BOW: pl. 2, f. 28]; [EBW]; [H&R, 111]; [MRS, 587]; [BRU2: 71, pl. 44, f. 25-33]; [Otakar Kudrna]
[cinxia-1u.jpg] 415x463 (~38Kb) upperside Finland, A: Lemland, Flaka, 6/1995, Photo © Tero Piirainen [cinxia-2v.jpg] 533x400 (~33Kb) underside A clearing in a pine forest at the village Nizhnii Tsasuchei, Onon District, SE Chita Province (Dahuria), S Transbaikalia, Siberia, Russia. 2nd July 1996, Photo © Oleg Kosterin [cinxia-3l.jpg] 550x371 (~54Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
[cinxia-3u.jpg] 595x400 (~58Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch

Larva on Plantago lanceolata, P. media [SPRK], (in Finland, Estonia) Veronica spicata, (in C.Europe) V. teucrium [Marko Nieminen]


M. c. cinxia
Morocco (Midle Atlas), W.Algeria Melitaea cinxia var. algirica Oberthür, 1915; Etud. Lépid. Comp., 10: 427, pl. 288, f. 2339 (preocc. Rühl, 1892), TL: Sebdou (Algeria)
Melitaea cinxia eupompe Hemming, 1933; Entomologist 66: 276 (repl. algirica Oberthür, 1915)
Melitaea cinxia cinxia ; [BMAT: 51, pl. 16, f. 22-27]

Larva on Centaurea pullata, Scorzonera pygmaea [BMAT], (in Spain) Plantago spp., (in Israel) Veronica macrostachya [BMAT]


eupompe
[eupompe-1v.jpg] 600x435 (~30Kb) underside Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre

M. c. clarissa Staudinger, 1901

Melitaea cinxia australis Verity, 1916
Melitaea cinxia narbonensis Fruhstorfer, 1917
Melitaea cinxia sicula Ragusa, 1920
 Melitaea cinxia deva Hemming, 1932
Melitaea cinxia castiliana Turati, 1920
Melitaea cinxia carpetana Verity, 1929
M. c. atlantis Le Cerf, 1928
Morocco (High Atlas, Toubkal Massif) Melitaea cinxia atlantis Le Cerf, 1928; Encycl. ent., Serie B, 3 (3): 224, TL: Dj. Tachdirt (High Atlas, Morocco)
Melitaea cinxia atlantis ; [H&R, 111]; [BMAT: 52, pl. 16, f. 28-32]
[atlantis-1u.jpg] 600x435 (~47Kb) upperside Morocco, Djebel Azourki, Haut Atlas, Photo © Jean Delacre [atlantis-2v.jpg] 600x435 (~28Kb) underside Morocco, Djebel Azourki, Haut Atlas, Photo © Jean Delacre

M. c. oasis Huang & Murayama
Melitaea cinxia oasis ; [MRS, 587]
M. c. tschujaca Seitz, 1908
Altai - E.Siberia Melitaea cinxia tschujaca Seitz, 1908; , TL: Altai
Melitaea cinxia arelatia Fruhstorfer, 1910
Melitaea cinxia tschujaca ; [BRU2, 71]
[tschujaca-1mu.jpg] 698x500 (~48Kb) upperside male Highland dwarf birch (Betula rotundifolia) tundras on a ledge of the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2300 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998, Photo © Oleg Kosterin

M. c. amardea Grum-Grshimailo, 1895
M. c. heynei Rühl, 1895
Alai Melitaea cinxia heynei Rühl, 1895; , TL: Alai
Melitaea cinxia heynei ; [BRU2, 71]
M. c. sacarina Fruhstorfer, 1917
cinxia sacarina Fruhstorfer, 1917; , TL: Saratov
Melitaea cinxia sacarina ; [BRU2, 71]
M. c. karavajevi Obraztsov, 1936
Ukraine cinxia karavajevi Obraztsov, 1936; , TL: Zhurovka, Kherson Region, Ukraine
Melitaea cinxia karavajevi ; [BRU2, 71]
Melitaea arcesia Bremer, 1861
S.Siberia (mountains), C.Yakutia, Magadan, Amurland, Primorye, Mongolia, N.China - C.China. See [About maps] Blackvein Fritillary
Melitaea arcesia Bremer, 1861; Mélanges biol. Acad. St.-Petersb. 3: 538, TL: Baical and Dahuria
?Melitaea arcesia Bremer, 1861; Bull. Acad. Imp. Sci. St. Petersb., 3: 466
Melitaea arcesia leechi Alphéraky, 1895
arcesia sikkimensis (Moore, 1901)
Melitaea arcesia irma Higgins, 1941
Melitaea arcesia, NSG Voucher Specimen [Wahlberg]
 Melitaea baicalensis ; Kosterin
Melitaea arcesia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name); [BOW: pl. 196, f. 5]; [BRU2: 71, pl. 41, f. 4-9]
Misidentified/misapplied:
Melitaea baicalensis Bremer, 1861 (= Mellicta sp.); Bremer, 1861, Mélanges biol. Acad. St.-Petersb. 3: pl. 1, f. 6
M. a. arcesia
Transbaikalia Melitaea arcesia arcesia ; [BRU2, 71]
M. a. minor Elwes, 1899
Altai Melitaea baicalensis minor ;
Melitaea arcesia minor ; [BRU2, 71]
[minor-1fv.jpg] 1060x700 (~83Kb) underside female An alpine meadow in the valley of a headwater of the Chikty Rivulet, 2500 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998 (on Tripleurospermum ambiguum), Photo © Oleg Kosterin [minor-1mu.jpg] 460x528 (~42Kb) upperside male An alpine meadow on a southern slope in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Tripleurospermum ambiguum), Photo © Oleg Kosterin [minor-1mv.jpg] 672x652 (~74Kb) underside male An alpine meadow on a southern slope in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Myosotis sp.), Photo © Oleg Kosterin

M. a. carmana Fruhstorfer, 1915
Sayan Melitaea carmana Fruhstorfer, 1915
dabanica Baranchikov, 1979
Melitaea arcesia carmana ; [BRU2, 71]
M. a. chuana Grum-Grshimailo, 1893
Melitaea arcesia chuana Grum-Grshimailo, 1893; , TL: Amdo
Melitaea arcesia chuana ; [BRU2, 71]
M. a. rucephala Fruhstorfer, 1915
Inn-Shan Mts. Melitaea arcesia rucephala ; [BRU2, 71]
Melitaea sindura Moore, 1865
Kashmir, Chitral, Tibet. See [About maps] Melitaea sindura Moore, 1865; Proc. zool. Soc. Lond. 1865 (2): 496
Melitaea sindura ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name); [BOW: pl. 195, f. 7]
Melitaea arcesia sindura ;
?M. s. honei Belter, 1942
Melitaea sindura honei Belter, 1942; , TL: NW.Yunnan
Melitaea balbita Moore, 1874
Pakistan, W.Himalayas. See [About maps] Melitaea balbita Moore, 1874; Proc. zool. Soc. Lond. 1874 (1): 268
Melitaea balbita ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea arcesia balbita ;
Melitaea amoenula C. & R. Felder, [1867]
Tibet. See [About maps] Melitaea amoenula C. & R. Felder, [1867]; Reise Fregatte Novara, Lep. Rhop. (3): 392
Melitaea arcesia amoenula Felder, 1867
Melitaea amoenula, NSG Voucher Specimen [Wahlberg]
 Melitaea amoenula ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)
Melitaea jezabel Oberthür, 1888
Tibet. See [About maps] Melitaea jezabel Oberthür, 1888; Etudes d'Ent. 11: 17
Melitaea jezabel Oberthür, ?1896
Melitaea jezabel yunnana Watkins, 1927; , TL: Shuantan, Loma valley, NW.Yunnan
= ; Huang, 2001, Neue Entomologische Nachrichten, 51: 103
Melitaea jezabel ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name); [MRS, 588]
Melitaea arcesia jezabel ;
Melitaea bellona Leech, [1892]
W.China. See [About maps] Melitaea bellona Leech, [1892]; Butts China Japan Corea (1): 219 (1892-1894)
Melitaea bellona, NSG Voucher Specimen [Wahlberg]
 Melitaea bellona ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name); [BOW: pl. 196, f. 3]; [MRS, 588]
Melitaea arcesia bellona ;
fiMelitaea diamina (Lang, 1789)
CEU, S.Europe, S.Siberia, NE.China, S.Ussuri, Korea, Japan. See [About maps] Tummaverkkoperhonen False-heath Fritillary Kovallnätfjäril Sotnätfjäril Silberscheckenfalter Doncella oscura Przeplatka diamina
Papilio diamina Lang, 1789; Verz. Schmett. Gegend. Augsburg (ed. 2): 44, TL: Augsburg
Melitaea dictynna alpestris Fruhstorfer, 1917
Melitaea diamina, NSG Voucher Specimen [Wahlberg]
Melitaea diamina, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Melitaea diamina, Svenska fjärilar [SVF]
Melitaea diamina, Norges Sommerfugler [NORLEP]
Melitaea diamina, Moths and Butterflies of Europe [MBE]
Melitaea diamina, Europaische Schmetterlinge [Mario Maier]
Melitaea diamina, Captain's European Butterfly Guide [Simon Coombes]
 Mellicta dictynna ; [MRS, 586]
Melitaea diamina ; [BOW: pl. 3, f. 1]; [EBW]; [H&R, 118]; [MRS, 588]; [BRU2: 75, pl. 47, f. 10-18]; [Otakar Kudrna]
[diamina-1u.jpg] 500x406 (~27Kb) upperside FINLAND: Orivesi, 1995, Photo © Juha Salokannel [diamina-2u.jpg] 500x412 (~32Kb) upperside Finland, Orivesi, Siitama, 4.7.1996, Photo © Tero Piirainen [diamina-3u.jpg] 500x419 (~26Kb) upperside Finland, Orivesi, Siitama, 4.7.1996, Photo © Tero Piirainen
[diamina-4.jpg] 500x595 (~64Kb) Finland, Orivesi, Siitama, 4.7.1996, Photo © Tero Piirainen

Larva on Valeriana officinalis, V. sambucifolia [SPRK], Veronica spp., Valeriana spp., Melampyrum spp. [MRS], Valeriana, Plantago, Veronica, Melampyrum, Patrinia, Polygonum [BRU2, 75]


M. d. diamina
Melitaea diamina diamina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name)
M. d. hebe (Borkhausen, 1793)
E.Europe - S.Siberia - Amur, Ussuri Papilio hebe Borkhausen, 1793
Papilio dictynna Esper, 1777
Papilio corythalia Hübner, [1799-1800]; Samml. eur. Schmett. [1] : pl. 3 (1799-1800), f. 15-16
orthia Hübner, 1818
lebedevi Obraztsov, 1936
Melitaea diamina hebe ; [BRU2, 75]
M. d. badukensis Alberti, 1969
Caucasus Major
M. d. vernetensis Rondou, 1902²
diamina vernetensis Rondou, 1902²; , TL: Vernet-les-Bain, E. Pyrenees
Melitaea diamina vernetensis ; [H&R, 119]
M. d. codinai Sagarra, 1932²
diamina codinai Sagarra, 1932²; , TL: Tortosa, NE.Spain
Melitaea diamina codinai ; [H&R, 119]
M. d. erycinides Staudinger, 1892
Melitaea diamina erycinides Staudinger, 1892; , TL: Kentei Mts.
Melitaea diamina erycinides ; [Oleg Kosterin]; [BRU2, 75 (?)]
[erycinides-1mu.jpg] 400x300 (~30Kb) upperside male A meadow at tiny aspen at a granite rocky ridge crest on a moderate elevation Adon-Chelon, at the mountain Tsagan-Obo, 10 km NNW of the village Tasyrkhoi, SE Chita region (Dahuria), Transbaikalia, Siberia, Russia. 9th July 1996, Photo © Oleg Kosterin

M. d. erycina (Lederer, 1853)
diamina erycina (Lederer, 1853); , TL: Altai
Mellicta dictynna erycina ; [MRS, 586]
Melitaea diamina erycina ; [BRU2, 75 (?)]
Melitaea protomedia Ménétriés, 1859
Amur, C.China, E.China, Korea, Japan. See [About maps] Melitaea protomedia Ménétriés, 1859; Bull. phys.-math. Acad. Sci. St. Pétersb., 17: 214, TL: [Amur region, Russia]
f. argentea Fixsen, 1887
Melitaea diamina protomedia ; [MRS, 588]
Melitaea protomedia ; [BOW: pl. 3, f. 1 (text only)]; [BRU2: 75, pl. 47, f. 19-21]

Larva on Veronica [BRU2, 76]

M. p. protomedia
Amur, Ussuri Melitaea protomedia protomedia ; [BRU2, 75]
M. p. regama Fruhstorfer, 1915
SW.China - Korea protomedia regama Fruhstorfer, 1915; , TL: Chang Yang, SW.China
Melitaea diamina regama ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [MRS, 588]
Melitaea proomedia regama ; [BRU2, 75]

The "minerva" species-group
"minerva" ; Achtelik, 1999, Atalanta 30 (1/4): 63
Melitaea minerva Staudinger, 1881
Tian-Shan. See [About maps] Melitaea minerva Staudinger, 1881; Stett. ent. Ztg 42: 289, TL: Ala Tau [Dzhungarsky]
Melitaea minerva, NSG Voucher Specimen [Wahlberg]
 Melitaea minerva ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [BRU2: 70, pl. 42, f. 1-15]
M. m. minerva
N.Tian-Shan, Dzhungarsky Alatau Mts. Melitaea minerva minerva ; [BRU2, 70]
M. m. pseudotersa Kolesnichenko & Churkin, 2000
Inner Tian-Shan (Baidulu Mts.) Melitaea minerva pseudotersa Kolesnichenko & Churkin, 2000; , TL: Inner Tian-Shan (Baidulu Mts.)
Melitaea minerva pseudotersa ; [BRU2, 70]
M. m. repens Kolesnichenko & Churkin, 2000
Kara-Tau Melitaea minerva repens Kolesnichenko & Churkin, 2000; , TL: Kara-Tau Mts.
Melitaea minerva repens ; [BRU2, 70]
M. m. tersa Kolesnichenko & Churkin, 2000
Karzhantau, Chatkal Melitaea minerva tersa ; [BRU2, 70]
M. m. acerba Kolesnichenko & Churkin, 2000
Chatkalsky Mts. (Lake Sary-Chelek) Melitaea minerva acerba Kolesnichenko & Churkin, 2000; , TL: Chatkalsky Mts. (Lake Sary-Chelek)
Melitaea minerva acerba ; [BRU2, 70]
Melitaea pallas Staudinger, 1886
-. See [About maps] Melitaea pallas Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 234, TL: [N. slope of the Alai Mts., Kirghizia]
Melitaea pallas, NSG Voucher Specimen [Wahlberg]
 Melitaea pallas ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 165 (name); [BOW: pl. 196, f. 6]; [MRS, 587]; [BRU2: 70, pl. 42, f. 16-30]
M. p. pallas
Alai, Transalai, Inner Tian-Shan Melitaea pallas pallas ; [BRU2, 70]