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Heterosais Godman & Salvin, [1880]
Heterosais Godman & Salvin, [1880]; Biol. centr.-amer., Lep. Rhop. 1: 60, TS: Ithomia nephele Bates
Rhadinoptera d'Almeida, 1922; Mélanges lép. 1: 71, TS: Ithomia nephele Bates
= ; [NL4A, 191]
Heterosais, NSG Voucher Specimen [Wahlberg]
Heterosais (Godyridini) ; [EcuL]; Lamas, 1999, Butterflies of the world, 3: 7; [NL4A, 191]
Heterosais (Godyridina) ; [Wahlberg]

Heterosais edessa (Hewitson, [1854])
Costa Rica - Bolivia. See [About maps] Ithomia edessa Hewitson, [1854]; Ill. exot. Butts 1 (12/13): [33] (1855), pl. [17], f. 42 ([1855]), TL: Brazil
Heterossais[sic] edessa ; Brown & Mielke, 1967, J. Lep. Soc. 21 (2): 93
Heterosais edessa ; [NL4A, #367]

Heterosais giulia (Hewitson, [1855])
Venezuela. See [About maps] Ithomia giulia Hewitson, [1855]; , TL: Colombia
Heterosais giulia ; [NL4A, #368]; [BOW: pl. 46, f. 2]
Colombia breyeri (Köhler, 1929) (Velamysta); , TL: "Argentina"
= ; [NL4A, #368a]
Heterosais giulia giulia ; [NL4A, #368a]
H. g. nephele (Bates, 1862)
Brazil(Amazonas) Ithomia nephele Bates, 1862; Trans. linn. Soc. Lond. 23 (3): 548, TL: Tabatinga
Hymenitis nephele ; Butler & Druce, 1874, Proc. Zool. Soc. Lond. 1874 (3): 333
Heterosais nephele ; Godman & Salvin, [1880], Biol. centr.-amer., Lep. Rhop. 1: 61; [BOW: pl. 46, f. 3]; Lamas, 1999, Butterflies of the world, 3: 7, pl. 16, f. 18
Heterosais nephele nephele ; [EcuL]
Heterosais edessa nephele ; [BCR, 237]
Heterosais giulia nephele ; [NL4A, #368d]
H. g. gedera (Hewitson, 1869)
Colombia giulia gedera (Hewitson, 1869) (Ithomia); , TL: [Colombia]
Heterosais giulia gedera ; [NL4A, #368c]
H. g. cadra (Godman & Salvin, 1878)
Panama Ithomia cadra Godman & Salvin, 1878; Ann. Mag. nat. Hist. (5) 2 (9): 259, TL: Panama
Ithomia cadra ; Godman & Salvin, [1880], Biol. centr.-amer., Lep. Rhop. 3: pl. 2, f. 5
Heterosais cadra ; Godman & Salvin, [1880], Biol. centr.-amer., Lep. Rhop. 1: 61
Heterosais giulia cadra ; [NL4A, #368b]
H. g. pallidula Haensch, 1903
Ecuador giulia pallidula Haensch, 1903; , TL: Ecuador, Palmar, Balzapamba
edessa covella d'Almeida, 1958; , TL: Ecuador, Manabi, Cojimies
= Heterosais nephele pallidula ; [EcuL]
= ; [NL4A, #368e]
Heterosais nephele pallidula ; [EcuL]; Lamas, 1999, Butterflies of the world, 3: pl. 16, f. 18
Heterosais giulia pallidula ; [NL4A, #368e]
Panama Heterosais giulia ssp. ; Lamas, MS, [NL4A, #368f]
Colombia Heterosais giulia ssp. ; Lamas, MS, [NL4A, #368g]
Colombia Heterosais giulia ssp. ; Lamas, MS, [NL4A, #368h]
Brazil(Amazonas) Heterosais giulia ssp. ; Lamas, MS, [NL4A, #368i]


5.2.2008 (2)

References:
[About maps]
Warning! The maps are automaticly generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BCR]
DeVries, Philip J.; The Butterflies of Costa Rica; Vol. I: Papilionidae, Pieridae, Nymphalidae
[BOW]
Lewis, H. L., 1974; Butterflies of the World; ISBN 0 245 52097 X
[EcuL]
Tommaso & Luigi Racheli, 2001; An annotated list of Ecuadorian Butterflies (Lepidoptera: Papilionidae, Pieridae, Nymphalidae); Fragmenta entomologica 33 (2): 213-380
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group


Some related literature:
Bates, 1862
Contributions to an Insect Fauna of the Amazon Valley. Heliconidae Trans. linn. Soc. Lond. 23 (3): 495-566, pl. 55-56
Brown & Mielke, 1967
Lepidoptera of the Central Brazil Plateau. I. Preliminary list of Rhopalocera J. Lep. Soc. 21 (2): 77-106
Butler & Druce, 1874
List of the butterflies of Costa Rica, with descriptions of new species Proc. Zool. Soc. Lond. 1874 (3): 330-370
Godman & Salvin, 1878
Description of new species of Rhopalocera from Central and South America Ann. Mag. nat. Hist. (5) 2 (9): 257-266
Godman & Salvin, [1880]
Biologia Centrali-Americana. Rhopalocera. Vol. 1. (1879-1886) Biol. centr.-amer., Lep. Rhop. 1: 1-487, 3: pl. 1-112
Hewitson, [1854]
Illustrations of new species of exoctic Butterflies selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson. 1. (1851-1856) Ill. exot. Butts 1 (1): [19-20],[67-68],[77-78], pl. [10],[34],[39] (1852), (2): [57-62], pl. [29-31] (1852), (3): [5-6],[21-22],[79-80], pl. [3],[11],[40] (1852), (4): [63-66],[81-82], pl. [32-33],[41] (1852), (5): [11-12],[113-114],[117-118], pl. [6],[57],[59] (1853), (6): [23-24],[69-70],[115-116], pl. [12],[35],[58] (1853), (7): [7-8],[95-96],[99-100], pl. [4],[48],[50] (1853), (8): [9-10],[101-102],[119- 120], pl. [5],[51],[60] (1853), (9): [13-14],[25-26],[83-84], pl. [7],[13],[42] (1854), (10): [87-90],[97-98], pl. [44-45],[49] (1854), (11): [15-16],[27-28], pl. [8],[14] (1854), (12/13): [29-36],[51-54],[103-104], pl. [15-18],[36-37],[52] (1855), (14): [37-40],[105-106], pl. [19-20],[53] (1855), (15): [41-44],[75-76], pl. [21-22],[38] (1855), (16): [1-2],[45-46],[85-86], pl. [1],[23],[43] (1855), (17): [3-4],[17-18], pl. [2],[9] (1856), (18): [47-50],[107-110], pl. [24-25],[54-55] (1856), (19): [51-52],[111-112], pl. [26],[56] (1856), (20): [53-56],[91-94],[121-124], pl. [27-28],[46-47] (1856)
Lamas, 1999
Nymphalidae II. Ithomiinae Butterflies of the world, 3: 17pp, 16pls


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Keep in mind that the taxononic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.