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Quadrus Lindsey, 1925
Quadrus Lindsey, 1925; Ann. ent. Soc. Amer. 18: 99, TS: Papilio cerialis Stoll
Cuadrus ; Parra, Vargas & Tabares, 2000, (missp.)
Quadrus (Pyrgini) ; [NL4A, 54]
Quadrus cerialis (Stoll, [1782])
Mexico - Bolivia, .?., Surinam, .... See [About maps] Papilio cerialis Stoll, [1782]; in Cramer, Uitl. Kapellen 4 (32-32): 213, pl. 392, f. N, O, TL: Surinam
Hesperia orcus Fabricius, 1793; , TL: "Indiis"
= ; [NL4A, #1013]
Pythonides cerberus Hübner, [1819]; (repl. name), TL: Surinam
f. majorinus (Draudt, 1922) (Pythonides); , TL: Bolivia
= ; [NL4A, #1013]
Quadrus cerialis, Listing of Mexican Butterflies [ILMB]
Quadrus cerialis, Butterflies of America [BOA]
 oreus ; Mabille, 1903, (missp.)
Pythonides cerialis ; Godman & Salvin, [1894], Biol. centr.-amer., Lep. Rhop. 2: 358, 3: pl. 82, f. 2; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 738
Quadrus cerealis[sic] ; Freeman, 1969, J. Lep. Soc. 23 (Suppl. 2): 22 (note, missp.); [BOW: pl. 86, f. 24]
cerebus ; Hayward, 1973, (missp.)
Quadrus ceriales[sic] ; Brown & Mielke, 1967, J. Lep. Soc. 21 (3): 162
Quadrus cerialis ; [NL4A, #1013]
Quadrus contubernalis (Mabille, 1883)
Mexico - Colombia, Brazil. See [About maps] Pythonides contubernalis Mabille, 1883; Bull. C. R. Soc. ent. Belg. 27: lxxvi [76], TL: Brazil,French Guinana
Quadrus contubernalis contubernalis, Listing of Mexican Butterflies [ILMB]
Quadrus contubernalis contubernalis, Butterflies of America [BOA]
 Pythonides contubernalis ; Godman & Salvin, [1894], Biol. centr.-amer., Lep. Rhop. 2: 359, 3: pl. 82, f. 3-5
Quadrus contubernalis ; [BOW: pl. 86, f. 25]; [NL4A, #1014]

Larva on Piper pseudolindenii Young, 1991, J. Lep. Soc. 45: 366

Q. c. contubernalis
Brazil, French Guinana Pythonides praxis Plötz, 1884; Jb. nassau. Ver. Naturk. 37 (1884): 7, TL: French Guinana
= Pythonides contubernalis ; Godman & Salvin, [1894], Biol. centr.-amer., Lep. Rhop. 2: 359
= ; [NL4A, #1014a]
Quadrus contubernalis contubernalis ; [NL4A, #1014a]
Q. c. anicius (Godman & Salvin, [1894])
Mexico, Guatemala, ... Pythonides anicius Godman & Salvin, [1894]; Biol. centr.-amer., Lep. Rhop. 2: 360, 3: pl. 82, f. 6-7, TL: Guatemala, Zapote
Quadrus anicius ;
Quadrus contubernalis anicius ; [NL4A, #1014b]
Quadrus deyrollei (Mabille, 1887)
-. See [About maps] Pythonides deyrollei Mabille, 1887; , TL: French Guinana
Quadrus deyrollei ; [BOW: pl. 86, f. 25 (text)]; [NL4A, #1015]
Q. d. deyrollei
French Guinana, ... deyrolli ; de Jong, 1983, (missp.)
Quadrus deyrollei deyrollei ; [NL4A, #1015a]
Q. d. porta Evans, 1953
Brazil(Amazonas) deyrollei porta Evans, 1953; , TL: Brazil, Amazonas
Quadrus deyrollei porta ; [NL4A, #1015b]
Q. d. portulana (Plötz, 1884)
deyrollei portulana (Plötz, 1884) (Pythonides); , TL: S.Amer.
Quadrus deyrollei portulana ; [NL4A, #1015c]
Quadrus fanda Evans, 1953
Brazil(Pará), .... See [About maps] Quadrus fanda Evans, 1953; , TL: Brazil, Pará
Quadrus fanda ; [NL4A, #1016]
Quadrus francesius Freeman, 1969
Mexico. See [About maps] Quadrus francesius Freeman, 1969; J. Lep. Soc. 23 (Suppl. 2): 21, pl. 7, f. 1-4, pl. 15, f. 3, TL: Santa Rosa, Comitán, Chiapas, Mexico
Quadrus francesius ; [NL4A, #1017]
Quadrus ineptus (Draudt, 1922)
Peru. See [About maps] Pythonides ineptus Draudt, 1922; , TL: Peru
Quadrus ineptus ; [NL4A, #1018]
Quadrus jacobus (Plötz, 1884)
Brazil(Rio de Janeiro, Distrito Federal). See [About maps] Tagiades jacobus Plötz, 1884; , TL: Brazil, Rio de Janeiro
Quadrus zolus Mielke, 1968; J. Lep. Soc. 22: 5, f. 5-6. 38-42, TL: Jardim Zoológico, Distrito Federal, Brazil
= ; [NL4A, #1019]
Quadrus jacobus ; [NL4A, #1019]
Quadrus lugubris (R. Felder, 1869)
Trinidad, Brazil, .... See [About maps] Leucochitonea lugubris R. Felder, 1869; Verh. zool.-bot. Ges. Wien. 19: 476, TL: Mexico, Veracruz
Gindanes obscurascens Mabille & Boullet, 1917
Quadrus lugubris, Listing of Mexican Butterflies [ILMB]
Quadrus lugubris lugubris, Butterflies of America [BOA]
Quadrus lugubris, Butterflies and Moths of North America [BAMONA]
 Pythonides lugubris ; Godman & Salvin, [1894], Biol. centr.-amer., Lep. Rhop. 2: 361, 3: pl. 82, f. 11-13
Quadrus lugubris ; [BOW: pl. 86, f. 26]; [NL4A, #1020]
[lugubris-1.jpg] 447x482 (~28Kb) Belize, Central America, Photo © John Shuey

Q. l. lugubris
Mexico, Venezuela Gindanes truncata var. obscurascens Mabille & Boullet, 1917; , TL: Mexico; Venezuela
= ; [NL4A, #1020a]
lagubris ; Kendall & McGuire, 1975, (missp.)
Quadrus lugubris lugubris ; [NL4A, #1020a]
Q. l. ophia (Butler, 1870)
Venezuela, Colombia Achlyodes ophia Butler, 1870; Trans. ent. Soc. Lond. 1870 (4): 517, TL: Venezuela
Pterygospidea sephara Hewitson, 1878; Ann. Mag. nat. Hist. (5) 1 (4): 344, TL: "Brazil"
= ; [NL4A, #1020b]
Pythonides cobarus Möschler, 1879; Verh. zool.-bot. Ges. Wien 28 (Abh.): 211, TL: Colombia
= ; [NL4A, #1020b]
Udramonia ambla Plötz, 1882; Berl. ent. Z. 26 (2): 266, TL: S.Amer.
= ; [NL4A, #1020b]
Quadrus lugubris ophia ; [NL4A, #1020b]
Quadrus tros Evans, 1953
Brazil(Espírito Santo). See [About maps] Quadrus tros Evans, 1953; , TL: Brazil, Espírito Santo
Quadrus tros ; [NL4A, #1021]
Quadrus truncata (Hewitson, 1870)
Bolivia, Ecuador. See [About maps] Pterygospidea truncata Hewitson, 1870; , TL: Ecuador
Quadrus truncata ; [BOW: pl. 86, f. 26 (text)]; [NL4A, #1022]
Quadrus u-lucida (Plötz, 1884)
-. See [About maps] Pythonides u-lucida Plötz, 1884; , TL: Colombia
Quadrus u-lucida ; [NL4A, #1023]
Q. u. u-lucida
Colombia, ... lucida ; Mabille & Boullet, 1912, (missp.)
Quadrus u-lucida u-lucida ; [NL4A, #1023a]
Q. u. mimus (Mabille & Boullet, 1917)
Brazil(Rio Grande do Sul, Săo Paulo, Paraná), Paraguay, Argentina(Misiones) u-lucida mimus (Mabille & Boullet, 1917) (Pythonides); , TL: Brazil, Rio Grande do Sul; Paraguay
Quadrus u-lucida mimus ; [NL4A, #1023b]
Q. u. parabus Mielke, 1968
Brazil(Minas Gerais) Quadrus u-lucida parabus Mielke, 1968; J. Lep. Soc. 22: 5, f. 7-8, TL: Paraopeab woods, Minas Gerais, Brazil
Quadrus u-lucida parabus ; [NL4A, #1023c]

25.6.2007 (11)

References:
[About maps]
Warning! The maps are automaticly generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Paul Opler, Harry Pavulaan, Ray Stanford, Michael Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOW]
Lewis, H. L., 1974; Butterflies of the World; ISBN 0 245 52097 X
[ILMB]
Interactive Listing of Mexican Butterflies
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[John Shuey]
e-mail: jshuey at tnc.org; Director of Conservation Science; Indiana Office of The Nature Conservancy; http://www.nature.org
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea

Some related literature:
Brown & Mielke, 1967
Lepidoptera of the Central Brazil Plateau. I. Preliminary list of Rhopalocera (continued) J. Lep. Soc. 21 (3): 145-168
Butler, 1870
Descriptions of some new diurnal Lepidoptera, chiefly Hesperiidae Trans. ent. Soc. Lond. 1870 (4): 485-520
R. Felder, 1869
Diagnosen neuer von dem K. K. Oberlieutenant H. v. Hedemann in Mexico in den Jahren 1865-1867 gesammelter Lepidopteren. Erste Folge Verh. zool.-bot. Ges. Wien. 19: 465-480
Freeman, 1969
Records, new species, and new genus of Hesperiidae from Mexico J. Lep. Soc. 23 (Suppl. 2): 62pp
Godman & Salvin, [1894]; Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Hewitson, 1878
Descriptions of twenty new Species of Hesperidae from his own Collection Ann. Mag. nat. Hist. (5) 1 (4): 340-348
Mielke, 1968
Lepidoptera of the Central Brazil Plateau. II. New Genera, Species and Subspecies of Hesperiidae J. Lep. Soc. 22: 1-20
Möschler, 1879
Neue exotische Hesperidae Verh. zool.-bot. Ges. Wien 28 (Abh.): 203-230
Plötz, 1882
Einige Hesperiinen -- Gattungen und deren Arten Berl. ent. Z. 26 (2): 253-266
Plötz, 1884
Die Hesperiinen -- Gruppe der Achlyoden Jb. nassau. Ver. Naturk. 37 (1884): 1-55
Stoll, [1782]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Young, 1991
Notes on the natural history of Quadrus (Pythonides) contuberalis (Hesperiidae) in Costa Rica J. Lep. Soc. 45: 366-371

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxononic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.